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Plants were among the earliest organis ms in the fossil record, but they did not colonize land until about 450 million years ago, or in the last 10% of the age of the earth . The delay was probably caused by the complexities of obtaining water in an inherently dry environment. Roots and an advanced vascular system were necessary to gather water while an epidermis and stomata were necessary to conserve it. Water had to move along a transport path consisting of many tissue types. Some tissues such as xylem were modified for water movement whereas others such as parenchyma were not. This complexity has caused researchers to view the system as a black box, at least initially, and approach the subject by measuring force s and determining the resulting water flows without knowing the precise nature of the flow path. Conceptually, this is no great problem, but it leaves open the possibility for variation between ' resent add re ss: Department o f S oil and C r o p S c P iences, Texas A&M Univer s ity, C o l lege Station, Texas 77843 0066-4294/85 /060 1 -0473 $02 .00 BOYER species adapted to different
Annual Review of Plant Biology – Annual Reviews
Published: Jun 1, 1985
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