ISSN 1067-4136, Russian Journal of Ecology, 2017, Vol. 48, No. 4, pp. 326–334. © Pleiades Publishing, Ltd., 2017.
Original Russian Text © E.G. Lapteva, N.E. Zaretskaya, P.A. Kosintsev, E.L. Lychagina, A.V. Chernov, 2017, published in Ekologiya, 2017, No. 4, pp. 267–276.
First Data on the Middle to Late Holocene Dynamics of Vegetation
in the Upper Kama Region
E. G. Lapteva
*, N. E. Zaretskaya
, P. A. Kosintsev
, E. L. Lychagina
, and A. V. Chernov
Institute of Plant and Animal Ecology, Ural Brach, Russian Academy of Sciences, Yekaterinburg, 620144 Russia
Geological Institute, Russian Academy of Sciences, Moscow, 119017 Russia
Perm State Humanitarian Pedagogical University, Perm, 614990 Russia
Moscow State University, Moscow, 119991 Russia
Received July 6, 2016
Abstract⎯A detailed palynological record and the results of radiocarbon dating of sediments from the
Dedyukhinskii floodplain massif in the vicinity of Lake Chashkinskoe (the Upper Kama region; 59°23′ N,
56°34.5′ E) have been used to reconstruct basic stages in the Middle to Late Holocene dynamics of vegeta-
tion. The results show that in the Atlantic period broadleaf tree species played a secondary role in forest for-
mations of taiga and subtaiga types. Broadleaf–conifer forests became dominant in the Subboreal period,
with fir widely spreading in the forests during its second half. During the Subatlantic period, forest formations
acquired their recent taiga character.
Keywords: palynological analysis, radiocarbon dating, vegetation, mammal fauna, archaeological sites, Mid-
dle and Late Holocene, Upper Kama region
Studies on reconstructing paleoecological condi-
tions in the Upper Kama region have been initiated
only in recent years [1, 2]. On the other hand, numer-
ous archaeological sites have been investigated there
, and paleoecological data are therefore important
for reconstructing the environment of ancient man.
In the territory of Perm Krai, only fragmentary pal-
ynological data supported by the results of radiocar-
bon analysis have been obtained to date [4–6]. Paly-
nological research in the Upper Kama region, in the
vicinity of Lake Chashkinskoe, began with studies on
cultural layers in archaeological sites Lake Chash-
kinskoe IV and Lake Chashkinskoe II, which provided
a basis for characterizing plant paleocommunities of
the Neolithic and Eneolithic periods [7, 8].
The purpose of this study was to integrate the initial
palynological data and the results of radiocarbon dat-
ing of organogenic deposits from Dedyukhinskii
Island (adjoining the Lake Chashkinskoe sites) in
order to reconstruct the paleovegetation of the region
during the Middle and Late Holocene.
STUDY REGION, OBJECTS, AND METHODS
The swamped floodplain massif named Dedyukh-
inskii Island is near the cities of Solikamsk and
Berezniki, Perm Krai (59°23′ N, 56°34.5′ E). It con-
sists of seven different-aged floodplain generations
(Fig. 1) and is bordered on the right (in the down-
stream direction) by the Kama River channel opposite
of the village of Pyskor and on the left by Lake Chash-
kinskoe. This lake is actually a system of intercon-
nected oxbow lakes that separates the island from the
accumulative terrace of the Kama where all archaeo-
logical sites are concentrated .
The climate of this region is moderately continental,
with warm summers and cold winters: the January and
July isotherms of –16°С and 17.5°С pass near Berezniki.
Annual precipitation is about 600–700 mm .
The study region is near the boundary of middle
and southern taiga fir–spruce forests  that are
dominated by Picea obovata Ledeb. and Abies sibirica
Ledeb. and also include pines (Pinus sylvestris L. and
P. sibirica Du Tour) and larch (Larix sibirica Ledeb.).
Birches (Betula pendula Roth. and B. pubescens
Ehrh.), aspen (Populus tremula L.), and alder (Alnus
incana (L.) Moench.) form small-leaved forests, par-
ticularly along riverbanks and peatland margins, and
occur as an admixture in dark conifer stands. In south-
ern taiga forests, linden (Tilia cordata Mill.) grows in
the understory, and elms (Ulmus laevis Pall. and
U. glabra Huds.) often occur in river valleys. The
herb–dwarf shrub layer is composed by boreal and
nemoral species. The moss layer is thin and discontin-
uous. Today, secondary pine–birch and spruce–birch