Catalogue of the Tachinidae of Egypt (Diptera: Oestroidea)

Catalogue of the Tachinidae of Egypt (Diptera: Oestroidea) Tachinid flies are an important group of parasitoids in their larval stage, and all their hosts are of the Arthropoda, almost exclusively other insects, including important insect pests in agriculture and forestry. All known Egyptian taxa of the family Tachinidae are systematically catalogued. Synonymies, type localities, type depositories, world distributions by biogeographic realm(s) and country, Egyptian localities, and dates of collection are provided. A total of 72 tachinid species belonging to 42 genera, 15 tribes, and 4 subfamilies has been treated. Keywords: Tachinid flies, Egyptian taxa, World distribution, Egyptian localities, Dates of collection Background agriculture and forestry. They typically parasitize phytopha- Tachinidae are a large and cosmopolitan family of flies gous larvae of Lepidoptera and Coleoptera or nymphs of within the superfamily Oestroidea. It is the second largest Hemiptera and Orthoptera. Consequently, tachinid flies family in the order Diptera (Irwin et al. 2003), with some have been successfully applied in programs of biological 1500 recognized genera (O’Hara 2016) and more than control against different insect pests (Stireman et al. 2006; 8500 described species (O’Hara 2013) worldwide. How- O’Hara 2008 and Cerretti and Tschorsnig 2010). ever, the estimated true diversity of the family is probably No comprehensive taxonomic studies on the family double the number of the currently known species, mak- Tachinidae have been carried out in Egypt before. Only one ing this family a good candidate for the first largest family published study, Steyskal and El-Bialy (1967), listed the of Diptera (O’Hara, 2008 and 2013 and Pape et al. 2011). Egyptian species of dipterous families including Tachinidae Tachinid flies range in size from 2 to 20 mm, and most of based on literature and specimens deposited in the main them are larger than a house fly and noticeably more bris- Egyptian insect collections. However, despite its import- tly, especially on thoracic dorsum and third to fifth abdom- ance, the list included only the names of species within the inal segments; however, across the family there is a family ordered alphabetically without referring to their tremendous variety of shapes, colors, and degree of brist- suprageneric classification or to other important data such ling. In addition to features shared with other families of as original descriptions, type localities, synonymies, world the super family Oestroidea (Rhinophoridae, Sarcophagi- distribution, type depositories, and local distribution. An- dae, Calliphoridae, and Oestridae), adult tachinids can be otherstudy,anunpublished Ph.D.thesisbyDr. Fathy recognized by the developed postscutellum (a rounded Negm (Plant Protection Institute, Ministry of Agriculture) bulge below the hind part of the thoracic dorsum) (O’Hara was completed in 1987 and reviewed the taxonomy of 2008). The family is divided into four subfamilies: Dexiinae, Egyptian Tachinidae. The author mainly adopted the classi- Exoristinae, Phasiinae, and Tachininae. All four subfamilies fication of the Palaearctic catalogue of Herting (1984). are represented in Egypt as shown in the present results. All missing data and deficiencies in previous Egyptian Tachinid flies are nectar or honeydew feeders as adults studies have been perfected with an updated taxonomy and are effective pollinators (Tooker et al. 2006). However, of the Egyptian Tachinidae in the present catalogue. they are an important group of parasitoids in their larval The present study is one in a series of planned studies stage, and all their hosts are of the phylum Arthropoda, al- on different Egyptian dipteran families aiming to cata- most exclusively other insects, including insect pests in logue the order Diptera as a whole in Egypt. Egypt, as a part of the Great Desert Belt, is characterized Correspondence: elhawagry@gmail.com by a warm and almost rainless climate (El-Hawagry 2017). Entomology Department, Faculty of Science, Cairo University, Giza, Egypt © The Author(s). 2018 Open Access This article is distributed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted use, distribution, and reproduction in any medium, provided you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 2 of 19 Ecologists divide Egypt into eight ecological zones: Coastal and written in bold uppercase letters. Species-group Strip, Lower Nile Valley and Delta, Upper Nile Valley, headings are left-justified and written in bold italicized Fayoum, Eastern Desert, Western Desert, Sinai, and Gebel lowercase letters except the first letter which is written Elba (Fig. 1). These zones have a greater affiliation to the uppercase. Authorship of genera, subgenera, and species Palaearctic Region except for the Gebel Elba, the southeast- are written in lowercase letters except the first letter ern triangle of Egypt, which has a greater affiliation to the which is written in uppercase. Taxonomically valid Afrotropical Region (El-Hawagry and Gilbert 2014). genus-group names (senior synonyms) are listed again in bold italicized lowercase letters (except the first upper- Material and methods case letter) and left-justified under the headings followed The present catalogue is based mainly on specimens col- by the reference including author, year, and journal lected by the author and his co-workers from different pages. Taxonomically valid species-group names com- Egyptian localities, in addition to data compiled from bined with their original genera (senior synonyms) are specimens preserved in the main Egyptian insect collec- listed again in regular italicized lower case letters (except tions, especially EFC and PPDD. A great deal of infor- the first uppercase letter) and left-justified under the mation, including synonymies and distributional data is headings followed by the reference including author, obtained from relevant literature and website databases year, and journal pages. Type species for all nomenclatu- as well. Of the most important literature reviewed are rally available genus-group names are given after the ref- Kugler (1980a); Herting (1984); Cerretti and Freidberg erence line, followed by method of their fixation. For 2009;O’Hara et al. (2009); O’Hara and Cerretti (2016); each genus and species-group name, associated syno- and Pape and Thompson (2017). nyms are listed in a chronological order. They are writ- ten in regular italicized lowercase letters (except the first Scope uppercase letter), followed by the reference and other This catalogue treats all names of taxa of the family data as in senior taxa. The rank of types that currently Tachinidae recorded from Egypt. exist (e.g., syntype, holotype, lectotype) is given after the reference of each species-group taxon, followed by its Arrangement of taxa location (depository) in museums, which is given as All taxa are arranged alphabetically. Old world syno- abbreviations in parenthesis. The type locality of each nyms, including all available and unavailable names, are species-group taxon is given at the end after its type chronologically listed. depository. Countries and islands of type localities are usually broken down to states, provinces, archipelagos, Typographical treatment of names ecological zones, towns, and villages. These “sublocal- Family-group headings are centered and written in bold ities” are placed in parentheses after the main locality, e. uppercase letters. Genus-group headings are left-justified g., “Egypt (Sinai)” . The sublocalities may be more broken down to “smaller sublocalities”. These “smaller sublocalities” are written after a colon following the sub- locality, e.g., “Egypt (Sinai: Tor)”. Distribution A complete world distribution by biogeographic realm(s) and country of each tachinid species, as is known from the litera- ture, is listed in alphabetical order. Distribution localities pre- ceded by a question mark (?) are considered questionable. Egyptian localities and dates of collection The Egyptian localities from which each tachinid species is collected and dates of collection are given to map a picture for the distribution and activity periods of the species in the different ecological zones of Egypt. Local- ities within each ecological zone are arranged alphabetic- ally and written after a colon following the ecological zone, e.g., “Coastal Strip: Mariout”. Spelling of locality Fig. 1 Map of Egypt showing the ecological zones (taken from El- names in Egypt almost follows El-Hawagry and Gilbert Hawagry and Gilbert 2014) (2014) and El-Hawagry (2015 and 2017). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 3 of 19 The catalogue Plagia hilfii sinaica Villeneuve in Hermann and This catalogue presents four tachinid subfamilies as rep- Villeneuve, 1909: 157. Holotype female (CNC). Egypt resented in Egypt, and the number of taxa treated in the (Sinai). catalogue can be summarized in table. Distribution: PA: Algeria, China, Egypt, S. Europe (Albania, Bosnia and Herzegovina, Bulgaria, Croatia, France, Greece, Italy, Malta, Monaco, Montenegro, Subfamily Tribes Genera Species Macedonia, San Marino, Serbia, Slovenia), Israel, Dexiinae 2 6 6 Kazakhstan,Morocco,Russia (W.Russia,E.Siberia), Exoristinae 5 21 37 Transcaucasia. Phasiinae 3 5 7 Egyptian localities: Sinai:? Tachininae 5 10 16 Dates of collection: April. Doubtful taxa 6 Genus PERISCEPSIA Gistel Total number 15 42 72 Scopolia Robineau-Desvoidy, 1830: 268 (junior homo- nym of Scopolia Hübner). Type species: Musca carbo- naria Panzer, 1798, by subsequent designation of Zetterstedt, 1844: 1239). Periscepsia Gistel, 1848:x(nomen novum for Scopolia Subfamily DEXIINAE Robineau-Desvoidy, 1830). Tribe DEXIINI Phoricheta Rondani, 1861a:8(nomen novum for Genus TRIXICEPS Villeneuve Scopolia Robineau-Desvoidy, 1830). Trixiceps Villeneuve, 1936: 329. Type species: Ramonda Robineau-Desvoidy, 1863a: 790. Type spe- Trixiceps aegyptiaca Villeneuve, 1936, by monotypy. cies: Ramonda fasciata Robineau-Desvoidy, 1863 Trixiceps magnipalpis (Bezzi, 1922) (= Tachina spathulata Fallén, 1820) by original Paraprosena magnipalpis Bezzi, 1922: 154. Type(s) not designation. located. Libya (Benghazi). Subgenus PERISCEPSIA Gistel —Nominotypical Trixiceps aegyptiaca Villeneuve, 1936: 329. Egypt subgenus (Ikingi Mariout). Periscepsia (Periscepsia) handlirschi (Brauer and Distribution: PA: Egypt, Europe, Libya. von Bergenstamm, 1891) Egyptian localities: Coastal Strip: Mariout, Mersa Phorichaeta handlirschii Brauer and Bergenstamm, Matruh. 1891: 52. Unspecified type(s) (NHMW). Italy (Trentino: Dates of collection: April to August. Trafoi). Note: The specific epithet was spelled handlirschii in Tribe VORIINI the original description but was subsequently changed to Genus HYPOVORIA Villeneuve handlirschi (O’Hara 2009). Hypovoria Villeneuve, 1913a: 510 (as subgenus of Distribution: PA: China, Egypt, Israel, Europe. Voria Robineau-Desvoidy 1830). Type species: Voria Egyptian localities: Coastal Strip: Burg El-Arab, Mariout. (Hypovoria) hilaris Villeneuve, 1913, by monotypy. Dates of collection: February to October. Hypovoria hilaris (Villeneuve, 1913) Genus UCLESIA Girschner Voria (Hypovoria) hilaris Villeneuve, 1913a: 510. Uclesia Girschner, 1901: 69. Type species: Uclesia Holotype female (CNC). Tunisia (Sfax). fumipennis Girschner, 1901, by monotypy. Distribution: PA: Canary Is., C. Asia (Kyrgyzstan, Uclesia petiolata (Villeneuve, 1929) Tajikistan, Turkmenistan, Uzbekistan), China, Egypt, Nanoplagia petiolata Villeneuve, 1929a: 45. Holotype S. Europe (Albania, Bosnia and Herzegovina, Bulgaria, female (?EFC, not located). Egypt (Wadi Rishrash). Croatia, France, Greece, Italy, Malta, Monaco, Distribution: PA: Algeria, Tunisia, Egypt, Russia, Montenegro, Macedonia, San Marino, Serbia, Tadzhikistan. Slovenia), W. Europe (Andorra, Portugal (including Egyptian localities: Eastern Desert: Wadi Rishrash. Azores, excluding Madeira), Spain), Israel, Mongolia, Dates of collection: March. Russia (E. Siberia), Tunisia, Transcaucasia. Genus WAGNERIA Robineau-Desvoidy Egyptian localities: Coastal Strip: El-Marg, Mariout. Wagneria Robineau-Desvoidy, 1830: 126. Type spe- Dates of collection: February to July. cies: Wagneria gagatea Robineau-Desvoidy, 1830,by Genus NANOPLAGIA Villeneuve monotypy. Nanoplagia Villeneuve, 1929a: 45. Type species: Wagneria cunctans (Meigen, 1824) Plagia hilfii Strobl, 1902, by original designation. Tachina cunctans Meigen, 1824: 419. Holotype male Nanoplagia sinaica (Villeneuve, 1909) (MNHN). Germany. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 4 of 19 Wagneria riedeli Villeneuve, 1937: 405. Germany Exorista aenescens Macquart, 1850: 409. France (Frankfurt). (Lestrem). Distribution: PA: Egypt, Europe, Israel. Exorista longicornis Perris, 1852: 206. France (Grandes Egyptian localities: Coastal Strip: El-Burg, Mariout, Landes). Nuzha. Eastern Desert: Suez Road. Lower Nile Valley Tachina bizonata Zetterstedt, 1859: 6119. Sweden and Delta: Dokki. Sinai: Ein Mouussa. (Skäne: Sjobo). Dates of collection: January to July. Distribution: AF: ?Malawi, Yemen. OR: Orien. China [see O’Hara and Cerretti 2016: 19]. PA: Canary Is., C. Asia (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), Subfamily EXORISTINAE Egypt, Europe, Israel, Japan, Korea, Kuril Is., Madeira, Tribe ERYCIINI Mongolia, Pal. China [see O’Hara and Cerretti 2016: 19], Genus AMELIBAEA Mesnil Russia, Transcaucasia. Amelibaea Mesnil, 1955: 454 (as subgenus of Egyptian localities: Lower Nile Valley and Delta: Abu Phebellia Robineau-Desvoidy, 1846). Type species: Par- Rawash, Koubba. exorista tultschensis Brauer and Bergenstamm, 1891,by Dates of collection: February to April. monotypy. Aplomya metallica (Wiedemann, 1824) Amelibaea ?signifera (Villeneuve, 1929) Tachina metallica Wiedemann, 1824: 46. Lectotype Exorista signifera Villeneuve, 1929a: 44. Holotype male male (ZMUC). India. [by fixation of Townsend (1933: (?EFC, not located). Egypt (Mariout). 470)]. Remarks: The identity of this species is not clear since Tachina nigriventris Wiedemann, 1824: 43. India. the male holotype is apparently lost (Cerretti and Tachina notata Wiedemann, 1830: 653. Sudan (Nubia Freidberg 2009). Region [as “Nubien”]). Distribution: PA: Egypt, Israel (Cerretti and Freidberg Tachina socia Wiedemann, 1830: 654. Sudan (Nubia 2009). Region [as “Nubien”]). Egyptian localities: Coastal Strip: Mariout. Phorocera eucalypta Loew, 1852: 659. Mozambique Dates of collection: January. (Tete [as “Tette”]). Genus APLOMYA Robineau-Desvoidy Parexorista laeviventris Wulp, 1893: 173. Indonesia Aplomya Robineau-Desvoidy, 1830: 184. Type species: (Jawa). Aplomya zonata Robineau-Desvoidy, 1830 (= Tachina Tricholyga impexa Villeneuve, 1916: 494. D.R. Congo confinis Fallén, 1820), by subsequent designation of and South Africa (Eastern Cape). Robineau-Desvoidy (1863a: 459, 460). Distribution: AF: Widespread. OR: Widespread. Prozenillia Villeneuve, 1916: 487. Type species: Palaearctic: Egypt, Israel, Japan, Pal. China [see O’Hara Prozenillia distans Villeneuve, 1916, by monotypy. and Cerretti 2016: 19]. Wiedemanniomyia Townsend, 1933: 469. Type Egyptian localities: Upper Nile Valley: Asswan. species: Tachina metallica Wiedemann, 1824,by Dates of collection: Unknown. original designation. Genus CADURCIELLA Villeneuve Aplomyiella Mesnil, 1939: 31. Type species: Tricholyga Cadurciella Villeneuve, 1927: 120. Type species: impexa Villeneuve, 1916 (= Tachina metallica Cadurciella rufipalpis Villeneuve, 1927, by monotypy. Wiedemann, 1824), by original designation. Cadurciella tritaeniata (Rondani, 1859) Atricholyga Villeneuve, 1939: 9. Type species: Exorista tritaeniata Rondani, 1859: 147. Unspecified Tricholyga impexa Villeneuve, 1916 (= Tachina type(?MZF). Italy (Parma). metallica Wiedemann, 1824), by monotypy. Distribution: PA: Egypt, Europe (northwards to Aplomya confinis (Fallén, 1820) Scotland, S. Norway, Sweden, Japan. Tachina confinis Fallén, 1820b: 32. Syntypes, males Egyptian localities and dates of collection are not precisely and females (NHRS and/or MZLU). Sweden (Gotland). known. Recorded by Steyskal and El-Bialy (1967: 48). Phryxe zonata Robineau-Desvoidy, 1830: 159. France. Genus CESTONIA Rondani Phryxe servillei Robineau-Desvoidy, 1830: 159. France Cestonia Rondani, 1861a: 105. Type species: Cestonia (Paris). cineraria Rondani, 1861, by monotypy. Aplomya nitens Robineau-Desvoidy, 1830: 185. France Cestonia cineraria Rondani, 1861 (Yonne: St. Sauveur). Cestonia cineraria Rondani, 1861a: 106. Syntypes, Aplomya zonata Robineau-Desvoidy, 1830: 185. France males and females (MZF). Italy (Parma). (Paris). Distribution: PA: Egypt, Europe (Mediterranean, Tachina bicingulata Zetterstedt, 1844: 1133. Sweden northwards to S. France, Switzerland, and Italy), (Skäne: Esperod). Israel. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 5 of 19 Egyptian localities: Coastal Strip: Nuzha. Eastern Egyptian localities and dates of collection are not Desert: Wadi Hoff. Lower Nile Valley and Delta: Ezbet precisely known. El-Nakhl. Sinai: Wadi El-Tih. Drino (Palexorista) inconspicua (Meigen, 1830) Dates of collection: March to July. Tachina inconspicua Meigen, 1830: 369. Unspecified Cestonia rutilans Villeneuve, 1929 types, females (MNHN). Germany (Berlin). Cestonia rutilans Villeneuve, 1929b: 102. Syntypes, 1 Tachina bimaculata Hartig, 1838: 286. Germany. male and 1 female, not located. Egypt (Cairo). Tachina flavoscutellata Zetterstedt, 1844: 1042. Sweden. Distribution: AF: Senegal, Yemen. PA: Egypt, Libya. Masicera cursitans Rondani, 1861a: 21. Italy (Parma). Egyptian localities and dates of collection are not Sturmia lophyri Robineau-Desvoidy, 1863b: 892. precisely known. Germany. Genus CESTONIONERVA Villeneuve Drino inusta Mesnil, 1949: 25. Poland. Cestonionerva Villeneuve, 1929a: 43. Type species: Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Conogaster petiolata Villeneuve, 1910, by subsequent Turkmenistan, Uzbekistan), China, Egypt, Europe, designation of Townsend (1936: 137). Russia (W. Russia, W. Siberia), Taiwan, Transcaucasia. Cestonionerva petiolata (Villeneuve, 1910) Egyptian localities and dates of collection are not Conogaster petiolata Villeneuve in Becker, 1910: 144. precisely known. Holotype female (NHMW, not located). Yemen (Socotra). Subgenus ZYGOBOTHRIA Mik Cestonionerva proxima Villeneuve, 1929a: 43. Egypt Zygobothria Mik, 1891: 193. Type species: Sturmia (Dahshour). atropivora Robineau-Desvoidy, 1830, by original Distribution: AF: U.A. Emirates, Yemen. PA: Canary designation. Is., C. Asia (Kyrgyzstan, Tajikistan, Turkmenistan, Formosodoria Townsend, 1933: 475. Type species: Uzbekistan), Israel, Mongolia, North Africa, Pal. China Sturmia (Argyrophylax) dilabida Villeneuve, 1916 (= [see O’Hara and Cerretti 2016: 19]. Meigenia ciliata van der Wulp, 1881), by original Egyptian localities: Lower Nile Valley and Delta: Dahshour. designation. Dates of collection: April. Drino (Zygobothria) atropivora (Robineau-Desvoidy, Genus DRINO Robineau-Desvoidy 1830) Drino Robineau-Desvoidy, 1863a: 250. Type species: Sturmia atropivora Robineau-Desvoidy, 1830: 171. Drino volucris Robineau-Desvoidy, 1863 (= Tachina lota Syntypes, males and females (MNHN). France. Meigen, 1824), by original designation. Sturmia masakensis Curran, 1927c: 117. Uganda Subgenus PALEXORISTA Townsend (Masaka). Palexorista Townsend, 1921: 134. Type species: Distribution: AF: Widespread. AU: Australia. OR: Tachina succini Giebel, 1862 (as “Tichina succini India, Indonesia, Laos, Malaysia, Orien. China [see Giebel”), by monotypy. O’Hara and Cerretti, 2016: 19], Ryukyu Is., Sri Lanka. Prosturmina Mesnil, 1970b: 110 (as subgenus of Drino PA: Algeria,CanaryIs.,C.Asia(Kyrgyzstan, Robineau-Desvoidy). Type species: Sturmia vigilans Tajikistan, Turkmenistan, Uzbekistan), Egypt, all Villeneuve, 1933 (= Sturmia pulchra Curran, 1927), Europe (exceptBritishIs.,Scand.),Israel,Japan,Pal. by original designation. China[seeO’Hara and Cerretti 2016: 19], Morocco, Drino (Palexorista) gilva (Hartig, 1838) Russia, Transcaucasia, Tunisia. Tachina gilva Hartig, 1838: 288. Syntypes, males and Egyptian localities: Coastal Strip: Ramleh. Eastern females, not located. Germany. Desert: Ismailia, Suez Road. Lower Nile Valley and Drino aurora Mesnil, 1949: 15. Japan. Delta: El-Qubba, Dekernes, Maadi, Shubra. Distribution: PA: Egypt, Europe, Japan, Transcaucasia. Dates of collection: April to July. Egyptian localities and dates of collection are not Genus THECOCARCELIA Townsend precisely known. Recorded from Egypt by Steyskal and Thecocarcelia Townsend, 1933: 471. Type species: El-Bialy (1967: 48). Argyrophylax pelmatoprocta Brauer and Bergenstamm, Drino (Palexorista) imberbis (Wiedemann, 1830) 1891 (= Masicera acutangulata Macquart, 1851), by Tachina imberbis Wiedemann, 1830: 317. Syntypes, original designation. males (ZMUH, lost). Egypt. Thelycarcelia Townsend, 1933: 475. Type species: Sturmia zonata Curran, 1927b: 336. Uganda (Entebbe). Thelycarcelia thrix Townsend, 1933 (= Sturmia Distribution: AF: D.R. Congo, Kenya, Malawi, South sumatrana Baranov, 1932), by original designation. Africa, Sudan, Tanzania, U.A. Emirates, Uganda, Yemen. Thecocarcelia acutangulata (Macquart, 1851) PA: Canary Is., C. Asia (Kyrgyzstan, Tajikistan, Masicera acutangulata Macquart, 1851a: 478. Turkmenistan, Uzbekistan), Egypt, Iran, Israel, North Unspecified type, female (?MHNL). Switzerland (Chur Africa, Turkey, Turkmenia. [as “Coire”]). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 6 of 19 Masicera incedens Rondani, 1861a: 22. Italy (Parma). Robineau-Desvoidy, 1830 (= Tachina obliquata Fallén 1810), Argyrophylax pelmatoprocta Brauer and Bergenstamm, by subsequent designation of Robineau-Desvoidy (1863a:553). 1891: 344. Austria (Bisamberg: nr. Vienna). Chetogena Rondani, 1856: 68. Type species: Salia Distribution: AF: Widespread. PA: Palaearctic: Egypt, rondaniana Villeneuve, 1931. all Europe (except Turkey), Japan, Transcaucasia. Spoggosia Rondani, 1859: 182. Type species: Spoggosia Egyptian localities and dates of collection are not occlusa Rondani, 1859 (=Tachina obliquata Fallén 1810), precisely known. by monotypy. Stomatomyia Brauer and Bergenstamm, 1889: 98. Tribe ETHILLINI Type species: Chetogena filipalpis Rondani, 1859,by Genus ATYLOMYIA Brauer subsequent designation of Brauer (1893: 483). Atylomyia Brauer, 1898: 525. Type species: Atylomyia Chetogena aegyptiaca (Villeneuve, 1923) loewii Brauer, 1898, by monotypy. Tachinosalia aegyptiaca Villeneuve, 1923: 91. Holotype Atylomyia albifrons Villeneuve, 1911 male (EFC). Egypt. Atylomyia albifrons Villeneuve, 1911: 86. Lectotype Distribution: PA: Egypt, Israel. female (CNC). Egypt (Helwan). Egyptian localities: Coastal Strip: Mariout. Atylomyia rungsi Mesnil, 1962: 778. Holotype. Dates of collection: March. Morocco (Ait Melloul). Chetogena cercosa (Kugler, 1980) Distribution: PA: China, Egypt, Israel, W. Europe Chaetogena cercosa Kugler, 1980a: 31. Holotype male (Andorra, Portugal (including Azores, excluding (TAU). Israel (Elat). Madeira), Spain). Distribution: AF: U. A. Emirates. PA: Egypt, Israel. Egyptian localities: Lower Nile Valley and Delta: Egyptian localities: Sinai: Serabit El-Khadim. Helwan. Dates of collection: April. Dates of collection: January and February. Chetogena sinaica (Villeneuve, 1909) Stomatomyia acuminata var. sinaica Villeneuve in Tribe EXORISTINI Hermann and Villeneuve, 1909: 157. Holotype female Genus ALLOPROSOPAEA Villeneuve (CNC). Egypt (Sinai). Alloprosopaea Villeneuve, 1923: 89. Type species: Distribution: PA: Egypt, Israel. Alloprosopaea efflatouni Villeneuve, 1923, by monotypy. Egyptian localities: Sinai: South Sinai. Alloprosopaea efflatouni Villeneuve, 1923 Dates of collection: ?April. Alloprosopaea efflatouni Villeneuve, 1923: 90. Genus EXORISTA Meigen Unspecified type(s) (CNC). Egypt (Wadi Hoff). Exorista Meigen, 1803: 280. Type species: Musca Distribution: PA: Egypt, Israel. larvarum Linnaeus, 1758, by monotypy. Egyptian localities: Eastern Desert: Wadi Hoff. Lower Subgenus ADENIA Robineau-Desvoidy Nile Valley and Delta: Dahshour. Adenia Robineau-Desvoidy, 1863a: 1041. Type species: Dates of collection: April. Tachina grisea Robineau-Desvoidy, 1830 (= Tachina rustica Genus CHAETORIA Becker Fallén, 1810), by original designation. Chaetoria Becker, 1908: 113. Type species: Chaetoria Staegeria Robineau-Desvoidy, 1863a: 972 (junior homo- stylata Becker, 1908, by monotypy. nym of Staegeria Rondani 1856). Type species: Tachina Clistorrhinia Bezzi in Bezzi and Lamb, 1926: 570. pratensis Robineau-Desvoidy, 1830,byoriginaldesignation. Type species: Clistorrhinia aurifrons Bezzi, 1926,by Chaetotachina Brauer and Bergenstamm, 1889: 98. monotypy. Type species: Tachina rustica Fallén, 1810, by monotypy. Chaetoria stylata Becker, 1908 Exorista (Adenia) rustica (Fallén, 1810) Chaetoria stylata Becker, 1908: 114. Lectotype female Tachina rustica Fallén, 1810: 264. Lectotype male (ZMHB). Canary Islands (Tenerife). (NHRS). Sweden (Skåne). Distribution: AF: Botswana, Mozambique, Nigeria, Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Senegal, U.A. Emirates, Yemen. PA: Algeria, Canary Is., C. Turkmenistan, Uzbekistan), China, Egypt, all Europe, M. Asia (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), East, Mongolia, Russia, Taiwan, Transcaucasia. Egypt, Europe, Morocco, Tunisia. Egyptian localities: Eastern Desert: Wadi Hoff. Egyptian localities: Coastal Strip: Ramleh. Upper Nile Dates of collection: March to June. Valley: Luxur. Subgenus EXORISTA Meigen —Nominotypical Dates of collection: January to September. subgenus Genus CHETOGENA Rondani Exorista Meigen, 1803: 280. Type species: Musca Salia Robineau-Desvoidy, 1830: 108 (junior homonym larvarum Linnaeus, 1758 (as “Musca larvarum Fabr.”), of Salia Hübner, 1818). Type species: Salia echinura by monotypy. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 7 of 19 Exorista (Exorista) fasciata (Fallén, 1820) Distribution: PA: Egypt, Israel. Tachina fasciata Fallén, 1820a: 5. Syntypes, males and Egyptian localities: Sinai. females (NHRS and/or MZLU). Sweden (Skåne, Äsperöd Dates of collection: April. [as “Esperöd”] and Abusa). Exorista (Ptilotachina) xanthaspis (Wiedemann, 1830) Chetogena segregata Rondani, 1859: 181. Italy (Parma Tachina xanthaspis Wiedemann, 1830: 314. Syntypes, and Sicily); Malta. males and females (SMF). Sudan (Nubia Region [as Distribution: NE: Yukon, introduced and established in “Nubien”]). Ontario, Québec and New England. OR: India. PA: Tachina pyrrhocera Wiedemann, 1830: 314. Sudan China, Egypt, Europe, Japan, M. East, Mongolia, Russia, (Nubia Region [as “Nubien”]). Taiwan, Transcaucasia. Tachina fallax pseudofallax Villeneuve, 1920: 151. Egyptian localities: Upper Nile Valley: Assiout. South Africa (Eastern Cape). Dates of collection: March. Larvaevora (Ptilotachina) fallax aethiopica Exorista (Exorista) larvarum (Linnaeus, 1758) Rohdendorf, 1931: 348. Sudan (Wad Medani). Musca larvarum Linnaeus, 1758: 596. Unspecified Distribution: AF: Madagascar, Seychelles, Sudan, U.A. Type(s) (LSUK). Europe. Emirates, Yemen. AU: N. Australasian. OR: India, Tachina moreti Robineau-Desvoidy, 1853: 534. France Indonesia, Orien. China [see O’Hara and Cerretti 2016: (Auxerre). 19], Ryukyu Is., Taiwan. PA: C. Asia (Kyrgyzstan, Distribution: NE: New England, Ontario, Québec, Tajikistan, Turkmenistan, Uzbekistan), Egypt, all Europe Yukon. OR: India. PA: C. Asia (Kyrgyzstan, Tajikistan, (except British Is., Scand.), Israel, Mongolia, Pal. China Turkmenistan, Uzbekistan), China, all Europe, Japan, M. [see O’Hara and Cerretti 2016: 19], Russia, East, Mongolia, North Africa, Russia, Taiwan, Transcaucasia. Transcaucasia. Egyptian localities: Lower Nile Valley and Delta: Abu Egyptian localities: Coastal Strip: Alexandria, Mariout, Rawash. Sinai: St. Katherine. Nuzha. Eastern Desert: Ismailia. Lower Nile Valley and Dates of collection: February to August. Upper Nile Delta: Bahtim, Dessouk, Dokki, Embaba, Sakha, Tanta. Valley: Asswan. Fayoum: ?. Sinai: El-Arish. Upper Nile Valley: Sohag. Genus MACULOSALIA Mesnil Dates of collection: Throughout the year. Maculosalia Mesnil, 1946: 62 (as subgenus of Subgenus PODOTACHINA Brauer and Bergenstamm Spoggosia Rondani 1859). Type species: Deuterammobia Podotachina Brauer and Bergenstamm, 1891: 350. maculosa Villeneuve, 1909, by monotypy. Type species: Tachina sorbillans Wiedemann, 1830,by Maculosalia maculosa (Villeneuve, 1909) subsequent designation of Townsend (1916: 8). Deuterammobia maculosa Villeneuve, 1909: 80. Exorista (Podotachina) sorbillans (Wiedemann, 1830) ?Holotype (?MNHN). Egypt (Sinai). Tachina sorbillans Wiedemann, 1830: 311. Lectotype Distribution: PA: Egypt, Iran, Tunisia. male (NHMW). Canary Islands (Tenerife). Egyptian localities: Sinai: ? Distribution: AF: Cameroon, Canary Is., D.R. Congo, Dates of collection: Unknown. Kenya, Malawi, Nigeria, Sierra Leone, Uganda. OR: India, Indonesia, Nepal, Orien. China [see O’Hara and Tribe GONIINI Cerretti 2016: 19], Philippines, Ryukyu Is., Sri Lanka, Genus BLEPHARELLA Macquart Taiwan, Thailand, Vietnam. Australasian: Australia, N. Blepharella Macquart, 1851b: 176. Type species: Australasian. PA: C. Asia (Kyrgyzstan, Tajikistan, Blepharella lateralis Macquart, 1851, by original Turkmenistan, Uzbekistan), Egypt, Europe, Israel, Japan, designation. Mongolia, China. Blepharella versatilis (Villeneuve, 1910) Egyptian localities and dates of collection are not Sturmia versatilis Villeneuve, 1910a: 253. Unspecified precisely known. Recorded from Egypt by Steyskal and type(s), 1 male (CNC). D.R. Congo. El-Bialy (1967: 49). Distribution: AF: D.R. Congo, Malawi, Nigeria, Sudan. Subgenus PTILOTACHINA Brauer and PA: ?Egypt. Bergenstamm Note: Villeneuve (1913b: 29) recorded this species Ptilotachina Brauer and Bergenstamm, 1891:350. from Egypt, based on a male in BMNH, however this Type species: Exorista florentina Herting, 1975, by record needs confirmation (O’Hara and Cerretti 2016). fixation of O’Hara et al.(2009: 94) under Article 70.3.2 Genus CLEMELIS Robineau-Desvoidy of the Code (ICZN, 1999). Clemelis Robineau-Desvoidy, 1863a: 481. Type spe- Exorista (Ptilotachina) rutilans Mesnil, 1970 cies: Zenillia ciligera Robineau-Desvoidy, 1830 Exorista rutilans Mesnil, 1970a: 113. Holotype male (= Tachina pullata Meigen, 1824), by original (MNHN). Egypt (Sinai). designation. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 8 of 19 Clemelis apicalis Villeneuve, 1923 Kom Osheem. Lower Nile Valley and Delta: Abu Clemelis apicalis Villeneuve, 1923: 92. Holotype male Rawash, Borkash, El-Marg, Ezbet El-Nakhl, Kafr (?EFC, not located). Egypt [as “Lower Egypt”]. Hakim, Pyramids, Shubra, Tanta, Zaqaziq. Distribution: PA: Egypt. Dates of collection: February to September. Egyptian localities: Coastal Strip: Mariout. Gonia capitata (De Geer, 1776) Dates of collection: April. Musca capitata De Geer, 1776: 23. Syntypes, Genus GONIA Meigen unspecified number and sex (?NHRS). Sweden. Salmacia Meigen, 1800: 38. Name suppressed by Rhedia testacea Robineau-Desvoidy, 1830: 75. France ICZN (1963: 339). (Paris). Gonia Meigen, 1803: 280. Type species: Gonia Gonia trifaria Zeller, 1842: 841. Austria (Wien), bimaculata Wiedemann, 1819, by subsequent Poland (Reinerz and Szczecin). designation of Sabrosky and Arnaud Jr (1965: 1075). Gonia maritima Perris, 1847: 495. France. Reaumuria Robineau-Desvoidy, 1830: 79. Type spe- Distribution: PA: China, Egypt, Europe, Mongolia, cies: Musca capitata De Geer, 1776, by subsequent des- Russia (W. Russia, W. Siberia), Transcaucasia. ignation of Robineau-Desvoidy (1863a: 733). Egyptian localities: Common throughout the country, Pissemya Robineau-Desvoidy, 1851b: 318. Type spe- particularly in: Coastal Strip: El-Burg, Mariout. Fayoum: ?. East- cies: Gonia atra Meigen, 1826, by monotypy. ern Desert: Wadi Rishrash, Wadi Wirak. Lower Nile Valley Turanogonia Rohdendorf, 1924: 228. Type species: and Delta: Giza, Helwan, Kerdassa, Tanta. Sinai: El-Arish, Ein Turanogonia smimovi Rohdendorf, 1924 (= Gonia Gedeirat,Ein Moweileh,Gebel Moussa,WadiEl-Lega,Wadi chinensis Wiedemann, 1824), by monotypy. Firan. Western Desert: Siwa Oasis. Upper Nile Valley: Asswan. Asiogonia Rohdendorf, 1928: 98. Type species: Dates of collection: February to October. Asiogonia asiatica Rohdendorf, 1928, by monotypy. Gonia ornata Meigen, 1826 Chrysocerogonia Rohdendorf, 1928: 98 (as subgenus of Gonia ornata Meigen, 1826: 3. Syntypes, males Salmacia Meigen, 1800). Type species: Salmacia (MNHN). France (Lyon). (Chrysocerogonia) ussuriensis Rohdendorf, 1928,by Gonia vittata Meigen, 1826: 3. Germany. monotypy. Gonia nervosa Meigen, 1826: 4. Germany (Hamburg). Eremogonia Rohdendorf, 1928: 98 (as subgenus of Rhedia vernalis Robineau-Desvoidy, 1830: 74. France. Salmacia Meigen, 1800). Type species: Salmacia Rhedia diversa Robineau-Desvoidy, 1830: 77. France. (Eremogonia) desertorum Rohdendorf, 1928,by monotypy. Gonia lateralis Zeller, 1842: 843. Poland (Glogow). Gonia atra Meigen, 1826 Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Gonia atra Meigen, 1826: 7. Unspecified type(s) Turkmenistan, Uzbekistan), China, Egypt, Europe (all), M. (MNHN). France. East, Mongolia, Russia (W. Russia, W. Siberia, S. Far East), Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Transcaucasia. Turkmenistan, Uzbekistan), China, Egypt, Europe, Egyptian localities: Common throughout the country, Kazakhstan, Mongolia, Russia (Siberia), Transcaucasia. particularly in: Lower Nile Valley and Delta: Abu Egyptian localities: Fayoum:? . Lower Nile Valley and Delta: Rawash, Tanta. Upper Nile Valley: Qena. El-Marg, Ezbet El-Nakhl, Kafr Hakim, Kerdassa, Sentrees. Dates of collection: February to August. Dates of collection: February to October. Genus MENDELSSOHNIA Kugler Gonia bimaculata Wiedemann, 1819 Mendelssohnia Kugler 1971: 69. Type species: Gonia bimaculata Wiedemann, 1819: 25. Syntypes, Mendelssohnia sinaica Kugler, 1971, by original females (NHMW, ZMUC). South Africa (Western Cape: designation. Cape of Good Hope). Mendelssohnia sinaica Kugler, 1971 Gonia cilipeda Rondani, 1859: 35. Italy (Parma and Mendelssohnia sinaica Kugler, 1971: 70. Holotype Malta). male (TAU). Egypt (Sinai: Wadi El-Sheikh). Gonia incerta Bigot, 1888: 10. (Tunisia. Tunis). Distribution: Egypt, Israel. Salmacia cabrerae Mesnil, 1956: 523. Canary Is. Egyptian localities: Sinai: Wadi El Sheikh. (Hierro). Dates of collection: Unknown. Distribution: AF: Widespread (excl. W. AF). OR: Orien. Genus NEALSOMYIA Mesnil China[seeO’Hara and Cerretti 2016: 19]. PA: Canary Is., C. Nealsomyia Mesnil, 1939: 31. Type species: Exorista Asia (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), (Alsomyia) triseriella Villeneuve, 1929, by original Europe, Israel, Madeira, North Africa, Pal. China [see designation. O’Hara and Cerretti 2016: 19], Transcaucasia. Nealsomyia triseriella (Villeneuve, 1929) Egyptian localities: CoastalStrip: Abu Qir. Eastern Exorista triseriella Villeneuve, 1929c: 185. Holotype Desert: Gebel Asfar, Ismailia, Wadi Hoff. Fayoum: male (CNC). Egypt (Wadi Hoff: nr. Helwan). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 9 of 19 Distribution: PA: Egypt, Israel. Ramonella Kugler, 1980b:67(nomen novum for Egyptian localities: Eastern Desert: Wadi Hoff. Ramona Kugler, 1980). Dates of collection: April to August. Ramonella mesnili (Kugler, 1980) Genus PSEUDOGONIA Brauer and Bergenstamm Ramona mesnili Kugler, 1980a: 41. Holotype male Pseudogonia Brauer and Bergenstamm, 1889: 100. (TAU). Israel (Negev: Ramon). Type species: Gonia cinerascens Rondani, 1859 Distribution: AF: Yemen. PA: Canary Is., Egypt, (= Tachina rufifrons Wiedemann, 1830), by monotypy. Turkey, Israel. Gaediogonia Townsend, 1927: 71. Type species: Egyptian localities: Lower Nile Valley and Delta: El- Gaediogonia jacobsoni Townsend, 1927 (= Tachina Marg. Sinai: Wadi Taibe, Wadi Watir. rufifrons Wiedemann, 1830), by original designation. Dates of collection: May to September. Pseudogonia fasciata (Wiedemann, 1819) Gonia fasciata Wiedemann, 1819: 25. Syntypes, Tribe WINTHEMIINI females (ZMUC). South Africa (Western Cape: Cape of Genus NEMORILLA Rondani Good Hope). Nemorilla Rondani, 1856: 66. Type species: Tachina Rhedia capensis Robineau-Desvoidy, 1830: 77. South maculosa Meigen, 1824, by original designation. Africa (Western Cape: Cape of Good Hope). Nemorilla floralis (Fallén, 1810) Reaumuria lalandii Robineau-Desvoidy, 1830: 80. Tachina floralis Fallén, 1810: 287. Holotype male South Africa (Western Cape: Cape of Good Hope). (NHRS). Sweden. Distribution: AF: South Africa, Zimbabwe. PA: Tachina notabilis Meigen, 1824: 266. Sweden. Andorra, Canary Is., Egypt, Portugal (including Azores, Tachina angustipennis Meigen, 1824: 333. W. Europe. excluding Madeira), Spain. Tachina pabulina Meigen, 1824: 358. Germany Egyptian localities and dates of collection are not (Hamburg). precisely known. Recorded from Egypt by Steyskal and Exorista laticella Macquart, 1850: 373. France El-Bialy 1967: 49). (Lestrem). Pseudogonia rufifrons (Wiedemann, 1830) Exorista arrogans Macquart, 1850: 374. France Tachina rufifrons Wiedemann, 1830: 318. Lectotype (Lestrem). female (ZMUC). China. Tachina intersita Walker, 1853: 72. England. Latreillia lalandii Robineau-Desvoidy, 1830: l06. South Nemorilla amica Rondani, 1859: 102. Italy (Parma). Africa (Western Cape: Cape of Good Hope). Aubaea campestris Robineau-Desvoidy, 1863: 186. Gonia cinerascens Rondani, 1859: 34. Italy (Parma). France. Gonia munroi Curran, 1927b: 339. Tanzania. Pitthaea nebulosa Robineau-Desvoidy, 1863: 189. Gonia ritchiei Cuthbertson and Munro, 1941:109. France. Nomen nudum. Distribution: PA: Egypt, Europe, Russia, Japan, North Gonia munroi curran, 1927b: 339) Tanzania (Tanganyika) Korea. and South Africa. Egyptian localities: Lower Nile Valley and Delta: Distribution: AF: Widespread, including Cape Verde, Bahtim, Ezbet El-Nakhl. Sinai: El-Arish. Nigeria, South Africa, Tanzania, U.A. Emirates, Yemen. Dates of collection: April to June. AU: Australia, Hawaii, Melanesia, N. Australasian. OR: India, Indonesia, Malaysia, Myanmar, Orien. China [see Subfamily PHASIINAE O’Hara and Cerretti 2016: 19], Pakistan, Philippines, Tribe CYLINDROMYIINI Ryukyu Is., Taiwan, Thailand. PA: C. Asia (Kyrgyzstan, Genus BESSERIA Robineau-Desvoidy Tajikistan, Turkmenistan, Uzbekistan), Egypt, all Europe Besseria Robineau-Desvoidy, 1830: 232. Type species: (except British Is.), Japan, Kazakhstan, Korea, M. East Besseria reflexa Robineau-Desvoidy, 1830, by monotypy. (Israel), Mongolia, Pal. China [see O’Hara and Cerretti Apostrophus Loew, 1871: 310, 311. Type species: 2016: 19], Russia, Transcaucasia. Apostrophus suspectus Loew, 1871 (=Actia zonaria Egyptian localities: Coastal Strip: Mariout, Nuzha. Loew, 1847), by subsequent designation of Coquillett Eastern Desert: Wadi Hoff. Gebel Elba: Wadi Edeib. (1910: 509). Lower Nile Valley and Delta: Abu Rawash. Sinai: Wadi Apostrophusia Townsend, 1933: 454. Type species: El-Tih. Apostrophus anthophilus Loew, 1871, by original Dates of collection: January to July. designation. Genus RAMONELLA Kugler Besseria anthophila (Loew, 1871) Ramona Kugler, 1980a: 40 (junior homonym of Apostrophus anthophilus Loew, 1871: 310. Type(s), Ramona Casey, 1886). Type species: Ramona mesnili male(s), not located. Germany (Bavaria: Munich- Kugler, 1980, by original designation. Geiselgasteig). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 10 of 19 Melia forcipata Bigot, 1881: 368. France (Hautes- Dionomelia Kugler, 1978b: 346. Type species: Alpes: Queyras, Abries). Dionomelia hennigi Kugler, 1978, by original designation. Distribution: NE: Alberta, Northwest Territories. PA: Dionomelia hennigi Kugler, 1978 Egypt, Europe, Mongolia. Dionomelia hennigi Kugler, 1978b: 346. Holotype Egyptian localities: Coastal Strip: Dekhila, Mariout. female (TAU). Israel (Dead Sea area: Ein Boqeq). Dates of collection: June to August. Distribution: AF: U.A. Emirates. PA: Andorra, Egypt, Genus CYLINDROMYIA Meigen Israel, Portugal (including Azores, excluding Madeira), Cylindromyia Meigen, 1803: 279. Type species: Spain. Musca brassicaria Fabricius, 1775, by monotypy. Egyptian localities: Sinai. Ocyptera Latreille, 1804: 195. Type species: Musca Dates of collection: April to September. brassicaria Fabricius, 1775, by subsequent designation of Genus LEUCOSTOMA Meigen Curtis (1837: 629). Leucostoma Meigen, 1803: 279. Type species: Exogaster Rondani, 1856: 78. Type species: Exogaster Ocyptera simplex Fallén, 1815, by subsequent monotypy carinatus Rondani, 1856 (= Ocyptera rufifrons Loew, of Meigen (1824: 234). 1844), by original designation. Leucostoma engeddense Kugler, 1966 Ocypterula Rondani, 1856: 78. Type species: Ocyptera Leucostoma engeddense Kugler, 1966: 177. Holotype pusilla Meigen, 1824, by original designation. female (TAU). Israel (Dead Sea area: Ein Gedi). Plesiocyptera Brauer and Bergenstamm, 1893: 56. Type Distribution: AF: South Africa, U.A. Emirates. PA: species: Ocyptera bicolor Wiedemann, 1819,by Algeria, Canary Is., Egypt, Israel, Madeira, NW Africa, monotypy. Spain, Turkey. Conopisoma Speiser, 1910: 144. Type species: Egyptian localities: Lower Nile Valley and Delta: Minia Conopisoma miraculum Speiser, 1910, by original El-Qamh, Tanta. designation. Dates of collection: February to May. Formicocyptera Townsend, 1933: 451. Type species: Leucostoma obsidianum (Wiedemann, 1830) Ocyptera atrata Fabricius, 1805, by original designation. Tachina obsidiana Wiedemann, 1830: 341. Lectotype Cylindromyia intermedia (Meigen, 1824) female (SMF). Sudan (Nuba [as “Nubien”]). Ocyptera intermedia Meigen, 1824: 212. Holotype Leucostoma marismortui Kugler, 1966: 179. Israel male (MNHN). ?Europe. (Dead Sea area: Ein Gedi). Ocyptera scalaris Loew, 1844: 240. Austria (Vienna). Distribution: AF: Sudan, Yemen. PA: Canary Is., Egypt, Ocyptera excisa Loew, 1845: 171. Romania (Mehadia). Europe, Israel, Malta, Saudi Arabia. Ocyptera excisa var. rufiventris Strobl, 1906: 338. Spain Egyptian localities: Upper Egypt: Asswan. (Villa Rutis). Dates of collection: Unknown. Ocyptera reinigi Enderlein, 1934: 132. Tajikistan (Pamir: Karasu). Tribe PHASIINI Distribution: Nearctic: Widespread. PA: C. Asia Genus PHASIA Latreille (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), Phasia Latreille, 1804: 195. Type species: Conops China, Egypt, Europe, Iran, Israel, Mongolia, Russia, subcoleoptratus Linnaeus, 1767, by subsequent Tadzhikistan, Transcaucasia, Ukraine, Uzbekistan. monotypy of Latreille (1805: 379). Egyptian localities: Coastal Strip: Alexandria, Mariout, Alophora Robineau-Desvoidy, 1830: 293. Type species: Max, Ramleh. Upper Nile Valley: Beni Suef. Syrphus hemipterus Fabricius, 1794, by subsequent des- Dates of collection: April to September. ignation of Robineau-Desvoidy (1863b: 226, as “Thereva Cylindromyia rufipes (Meigen, 1824) hemiptera de Fabricius”). Ocyptera rufipes Meigen, 1824: 215. Lectotype male Hyalomya Robineau-Desvoidy, 1830: 298. Type spe- (MNHN). France. cies: Phasia semicinerea Meigen, 1824 (=Phasia pusilla Distribution: AF: U.A. Emirates. OR: India, Pakistan. Meigen, 1824), by subsequent designation of Westwood PA: Egypt, all Europe (except British Is., Scand.), (1840: 140). Pakistan, Russia (W. Russia), Saudi Arabia, Phorantha Rondani, 1861b: 209. Type species: Transcaucasia, Ukraine. Phorantha musciformis Rondani, 1861 (= Conops Egyptian localities: Coastal Strip: Alexandria. Lower subcoleoptratus Linnaeus, 1767), by monotypy. Nile Valley and Delta: Gebel Asfar, Kubba. Paralophora Girschner, 1887: 412 (as subgenus of Dates of collection: October to February. Alophora Robineau-Desvoidy, 1830). Type species: Phasia pusilla Meigen, 1824, by monotypy. Tribe LEUCOSTOMATINI Alophorella Townsend, 1912: 45. Type species: Genus DIONOMELIA Kugler Thereva obesa Fabricius, 1798, by original designation. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 11 of 19 Phasia pusilla Meigen, 1824 Tachinomima Brauer and Bergenstamm, 1891: 383. Phasia pusilla Meigen, 1824: 198. Lectotype male Type species: Tachinomima expetens Brauer and (CNC). Germany (Stolberg). Bergenstamm, 1891 (= Micropalpus longirostris Phasia semicinerea Meigen, 1824: 199. Germany Macquart, 1844), by monotypy. (Stolberg). Hecatoepalpus Townsend, 1933: 467. Type species: Hyalomya carbonaria Robineau-Desvoidy, 1830: 300. Micropalpus prohecate Speiser, 1910, by original France (St. Sauveur). designation. Hyalomya corinna Robineau-Desvoidy, 1830: 301. Micropalpinus Enderlein, 1937: 441. Type species: France. Micropalpus pallidus Jaennicke, 1867, by original Phasia vitripennis Zetterstedt, 1859: 6169. Sweden designation. (Skane). Gymmantia Enderlein, 1937: 441. Type species: Hyalomya chorea Robineau-Desvoidy, 1863: 266. Micropalpus alboscutellatus Speiser, 1910, by original France. designation. Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Linnaemya comta (Fallén, 1810) Turkmenistan, Uzbekistan), China, Egypt, Europe (all), Tachina comta Fallén, 1810: 277. Unspecified type(s), Israel, Japan, Kazakhstan, Mongolia, Russia, Transcaucasia. females (NHRS) Sweden. Egyptian localities and dates of collection are not Tachina compta Meigen, 1824: 262. Unjustified precisely known. emendation of comta Fallén. Tachina fulgens Meigen, 1824: 259. Austria, France Subfamily TACHININAE and Germany. Tribe ERNESTIINI Linnaemya heraclei Robineau-Desvoidy, 1830: 53. Genus EURITHIA Robineau-Desvoidy France (Paris). Erigone Robineau-Desvoidy, 1830: 65 (junior homo- Linnaemya analis Robineau-Desvoidy, 1830: 54. nym of Erigone Audouin, 1826). Type species: Erigone France (Angers). anthophila Robineau-Desvoidy, 1830, by subsequent Linnaemya aestivalis Robineau-Desvoidy, 1830: 54. designation of Townsend (1932: 42). France. Eurithia Robineau-Desvoidy, 1844: 24. Type species: Linnaemya borealis Robineau-Desvoidy, 1830: 54. Erigone puparum Robineau-Desvoidy, 1830 (= Tachina France. caesia Fallén, 1810), by monotypy. Bonnetia longipes Robineau-Desvoidy, 1830: 55. Eurithia castellana (Strobl, 1906) France. Erigone castellana Strobl, 1906: 338. Holotype male Bonnetia oenanthis Robineau-Desvoidy, 1830: 56. (NMBA). Spain (Madrid). Holotype may be lost, see France (Paris). O’Hara et al. (2009). Micropalpus humeralis Robineau-Desvoidy, 1863: 119. Distribution: PA: Albania, Bosnia and Herzegovina, New name for Bonnetia longipes Robineau-Desvoidy, 1830. Bulgaria, China, Croatia, Egypt, Greece, Israel, Distribution: AF: Sudan. NE: Widespread. OR: India, Montenegro, Macedonia, Serbia, Slovenia, Transcaucasia. Nepal. PA: C. Asia (Kyrgyzstan, Tajikistan, Egyptian localities: Coastal Strip: El-Burg, Mariout. Turkmenistan, Uzbekistan), China, Egypt, Europe (all), Eastern Desert: Ismailia. Fayoum: ?. Lower Nile Valley Kazakhstan, Israel, Mongolia, Russia (W. Russia, W. and Delta: Abu Rawash. Upper Nile Valley: Asswan. Siberia, E. Siberia, S. Far East), Taiwan, Transcaucasia. Dates of collection: January to October. Egyptian localities: Eastern Desert: Wadi Garawi, Wadi Genus LINNAEMYA Robineau-Desvoidy Rishrash. Gebel Elba: ?. Lower Nile Valley and Delta: Linnaemya Robineau-Desvoidy, 1830: 52. Type spe- Mansouriya. cies: Linnaemya silvestris Robineau-Desvoidy, 1830 Dates of collection: January to March. (= Tachina vulpina Fallén, 1810), by subsequent designa- Linnaemya latigena Kugler, 1977 tion of Robineau-Desvoidy (1863a: 131). Linnaemyia latigena Kugler, 1977: 3. Holotype male Bonellia Robineau-Desvoidy, 1830: 56 (junior homo- (TAU). Egypt (Sinai: Bir Gifgafa [as “Refidim”]). nym of Bonellia Rolando, 1822). Type species: Bonellia Distribution: AF: U.A. Emirates. PA: Egypt, Israel. tessellans Robineau-Desvoidy, 1830, by subsequent des- Egyptian localities: Sinai: Bir Gifgafa, Mitla, Wadi ignation of Townsend (1916: 6). Firan. Elachipalpus Rondani, 1850: 169. Type species: Dates of collection: April. Micropalpus longirostris Macquart, 1845 (junior primary Linnaemya petiolata Kugler, 1971 homonym of Micropalpus longirostris Macquart, 1844; Linnaemya petiolata Kugler, 1971: 80. Unspecified = Elachipalpus rondanii Townsend, 1916), by original type(s), male(s) (TAU). Egypt (South Sinai: Abu designation. Marawath). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 12 of 19 Distribution: PA: Egypt. Egyptian localities. Coastal Strip: Alexandria, Amrya. Egyptian localities: Sinai: Abu Marawath. Lower Nile Valley and Delta: Abu Rawash, Ain Shams, Dates of collection: April to August. Beni Meggon, Cairo, Dokki, El-Baragil, Ezbet El-Nakhl, Linnaemya soror Zimin, 1954 Giza, Kafr Hakim, Kerdassa, Mansouriya, Maadi, Nahya, Linnaemyia soror Zimin, 1954: 266. Holotype male Shubra, Sinai: El-Arish. Sinai: El-Arish. Upper Nile Val- (ZIN). Tajikistan (Gorno-Badakhshan: Khorugh). ley: Assiout, Asswan. Distribution: PA: Canary Is., C. Asia (Kyrgyzstan, Dates of collection. Throughout the year. Tajikistan, Turkmenistan, Uzbekistan), China, Egypt, W. Genus PLESINA Meigen Europe (Andorra, Portugal (including Azores, excluding Plesina Meigen, 1838: 214. Type species: Tachina Madeira), Spain), S. Europe (Albania, Bosnia and phalerata Meigen, 1824, by monotypy. Herzegovina, Bulgaria, Croatia, Greece, Montenegro, Xanthopetia Townsend, 1933: 452. Type species: Macedonia, Serbia, Slovenia), Israel, North Africa, Tachina fascipennis Wiedemann, 1830, by original Russia, Transcaucasia, Turkmenia, Uzbekistan. OR: designation. India, Nepal. Kugleria Verbeke, 1970: 299 (junior homonym of Egyptian localities: Pyramids. Kugleria Bouwman, 1938). Type species: Plesina Dates of collection: September. fascipennis claripennis Mesnil, by monotypy. Plesina claripennis Mesnil, 1953 Plesina claripennis Mesnil, 1953b: 166. Holotype (not Tribe GRAPHOGASTRINI located). Israel (Jerusalem). Genus PHYTOMYPTERA Rondani Distribution: Egypt, Israel. Phytomyptera Rondani, 1845: 32, 33. Type species: Egyptian localities: Sinai. Phytomyptera nitidiventris Rondani, 1845 (= Tachina Dates of collection: ? nigrina Meigen, 1824), by monotypy. Plesina deserticola Kugler, 1978 Elfia Robineau-Desvoidy, 1850: 190. Type species: Plesina deserticola Kugler, 1978a: 93. Holotype male Actia cingulata Robineau-Desvoidy, 1830, by subsequent (TAU). Egypt (Sinai: Wadi Shag). designation of Robineau-Desvoidy (1863a: 672). Distribution: PA: Egypt, Israel. Phytomyptera lacteipennis Villeneuve, 1934 Egyptian localities: Sinai: Wadi Shag (South Sinai). Phytomyptera lacteipennis Villeneuve, 1934b: 71. Dates of collection: ? Lectotype female (CNC). Egypt (Suez). Genus ROSSIMYIOPS Mesnil Distribution: AF: U.A. Emirates. PA: Egypt, Europe, Rossimyiops Mesnil, 1953a: 145. Type species: Israel, Libya, Mongolia, Russia (W. Russia). Rossimyiops whiteheadi Mesnil, 1953, by monotypy. Egyptian localities: Eastern Desert: Suez Road. Mesnilomyia Kugler, 1972: 103. Type species: Dates of collection: March. Mesnilomyia magnifica Kugler, 1972, by original designation. Tribe MINTHOINI Persedea Richter, 2001: 25. Type species: Persedea Genus MINTHO Robineau-Desvoidy exquisita Richter, 2001, by original designation. Mintho Robineau-Desvoidy, 1830: 216. Type species: Rossimyiops achilleae (Kugler, 1972) Musca compressa Fabricius, 1787, by subsequent desig- Mesnilomyia achilleae Kugler, 1972: 107. Holotype nation of Rondani (1856: 79). male (TAU). Israel (‘Arad). Mintho compressa (Fabricius, 1787) Distribution: PA: Egypt, Israel. Musca compressa Fabricius, 1787: 346. Unspecified Egyptian localities: Sinai: St. Catherine, Wadi El- type(s) (ZMUC). Spain. Arbaein, Wadi Shag. Mintho capensis Robineau-Desvoidy, 1830: 217. South Dates of collection: July. Africa (Western Cape: Cape of Good Hope). Rossimyiops magnificus (Kugler, 1972) Tachina isis Wiedemann, 1830: 304. Egypt. Mesnilomyia magnifica Kugler, 1972: 105. Holotype Ocyptera vittata Costa, 1844: 94. Italy (Cervinara and male (TAU). Israel (‘Arad). Guagnano). Distribution: PA: Egypt, Iraq, Israel. Sericocera algira Macquart, 1849: 480. Algeria. Egyptian localities: Sinai: Tarfat El-Kadarin, Wadi Dexia thala Walker, 1849: 845. Morocco (Tangier). Nazeb, Wadi Sa’al, Nueiba’. Dexia isapis Walker, 1849: 848. Egypt. Dates of collection: April to September. Distribution: AF: Widespread, including D.R. Congo, Eritrea, Kenya, Nigeria, Somalia, South Africa, Sudan, Tribe SIPHONINI Tanzania, Yemen. PA: Algeria, Canary Is., Egypt, Europe, Genus ACTIA Robineau-Desvoidy Israel, Libya, Morocco, Transcaucasia, Tunisia. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 13 of 19 Actia Robineau-Desvoidy, 1830: 85. Type species: Roe- Taiwan, Thailand. PA: Japan, Israel, North Africa, P. selia lamia Meigen, 1838, by designation under the plen- China. AU: Australia, Melanesia, Micronesia, N. ary powers of ICZN. Australasian. Gymnophthalma Lioy, 1864: 1341. Type species: Egyptian localities: Lower Nile Valley and Delta: Giza, Tachina crassicornis Meigen, 1824, by monotypy. Sidi Ghazi, Torah. Upper Nile Valley: Beni Morr (Assiout). Gymnopareia Brauer and Bergenstamm, 1889:103. Dates of collection: May to August. Type species: Tachina crassicornis Meigen, 1824,by Peribaea palaestina (Villeneuve, 1934) monotypy. Actia palaestina Villeneuve, 1934a: 57. Holotype Actia crassicornis (Meigen, 1824) female (SMNS). Israel, (Rehovot [as “Rehoboth”]). Tachina crassicornis Meigen, 1824: 351. Syntypes, Actia alipes Villeneuve, 1942: 134. Egypt (Asswan [as males and females (MNHN). Probably Germany (Stolberg). “Assuan”]). See O’Hara (2009). Distribution: AF: U.A. E., Yemen. PA: C. Asia Thryptocera bicolor Macquart, 1834: 312. France (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), (Lille). China, Egypt, Iran, Israel. Thryptocera flavipalpis Macquart, 1848: 135. Egyptian localities and dates of collection are not Switzerland (Ütliberg). precisely known. Thryptocera nigripalpis Robineau-Desvoidy, 1851: 182. Genus SIPHONA Meigen France. Crocuta Meigen, 1800: 39. Name suppressed by ICZN. Thryptocera palpalis Rondani, 1859: 14. Italy (Parma). Siphona Meigen, 1803: 281. Type species: Musca Thryptocera claripennis Robineau-Desvoidy, 1863: 716 geniculata De Geer, 1776, by designation under the France, plenary powers of ICZN. Thryptocera scutellaris Rondani, 1865: 195. Italy Subgenus SIPHONA Meigen, 1803 —Nominotypical (Parma). subgenus. Distribution: PA: China, Egypt, Europe (all), Japan, Siphona (Siphona) efflatouni Mesnil, 1960 Kazakhstan, Mongolia, Russia, Transcaucasia. Siphona efflatouni Mesnil, 1960: 188. Holotype female Egyptian localities: Coastal Strip: Alexandria. Eastern (CNC). Egypt (Mariout). Desert: Ismailia. Lower Nile Valley and Delta: Abu Distribution: PA: Egypt, Israel. Rawash, Dokki, Giza, Kerdassa. Sinai: El-Arish. Upper Egyptian localities: Coastal Strip: Mariout. Sinai: El-Arish. Nile Valley: Assiout. Dates of collection: March to August. Dates of collection: April to September. Genus PERIBAEA Robineau-Desvoidy Tribe TACHININI Herbstia Robineau-Desvoidy, 1851: 184 (junior homo- Genus PELETERIA Robineau-Desvoidy nym of Herbstia Edwards, 1834). Type species: Herbstia Peleteria Robineau-Desvoidy, 1830: 39. Type species: tibialis Robineau-Desvoidy, 1851, by monotypy. Peleteria abdominalis Robineau-Desvoidy, 1830, by sub- Peribaea Robineau-Desvoidy, 1863a: 720. Type spe- sequent designation of Coquillett (1910: 586). cies: Peribaea apicalis Robineau-Desvoidy, 1863 (= Herb- Cuphocera Macquart, 1845: 267. Type species: stia tibialis Robineau-Desvoidy, 1851), by subsequent Micropalpus ruficornis Macquart, 1835, by original designation of Coquillett (1910: 587). designation. Strobliomyia Townsend, 1926a: 31. Type species: Acuphocera Townsend, 1926b: 37. Type species: Tryptocera fissicornis Strobl, 1910 (as “Thryptocera Acuphocera sumatrensis Townsend, 1926 (= Tachina fissicornis”)(= Thryptocera setinervis Thomson, 1869), iavana Wiedemann, 1819), by original designation. by original designation. Pleropeleteria Villeneuve, 1916: 470. Type species: Peribaea orbata (Wiedemann, 1830) Dejeania (Pleropeleteria) peringueyi Villeneuve, 1916 Tachina orbata Wiedemann, 1830: 336. Neotype (= Tachina lithanthrax Wiedemann, 1830), by monotypy. female (BMNH). India (Assam: Azra). Peletieriana Mesnil, 1970a: 951 (as subgenus of Gymnopareia (Actia) aegyptia Villeneuve, 1913a: 508. Peleteria Robineau-Desvoidy, 1830,as “Peletieria”). Type Egypt (Qaliûb). species: Echinomyia rustica Karsch, 1886, by original Actia nigripes Curran, 1927a: 6. D.R. Congo (Bas- designation. Congo: Boma). Peleteria meridionalis (Robineau-Desvoidy, 1830) Strobliomyia sororcula Mesnil, 1954: 16. D.R. Congo Faurella meridionalis Robineau-Desvoidy, 1830: 41. (Nord-Kivu: Rutshuru). Type(s) not located. France. Distribution: AF: Widespread. OR: India, Indonesia, Echinomyia ruficeps Macquart, 1845: 263. Italy. Orien. China [see O’Hara and Cerretti 2016: 19], Peleteria vernalis Robineau-Desvoidy, 1863: 617. Malaysia, Myanmar, Philippines, Ryukyu Is., Sri Lanka, France (Hyeres). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 14 of 19 Peleteria pedemontana Robineau-Desvoidy, 1863: 619. Genus UCLESIA Girschner Italy. Peleteria algira Robineau-Desvoidy, 1863: 621. Algeria. – Uclesia petiolata (Villeneuve, 1929) Peletieria capitata Zimin, 1961: 265. Turkmenistan (Chandyr Valley). Genus WAGNERIA Robineau-Desvoidy Distribution: PA: Algeria, Egypt, Isael, Italy, Kirghizia, Russia, Tadzhikistan, Transcaucasia, Uzbekistan. – Wagneria cunctans (Meigen, 1824) Egyptian localities: Coastal Strip: Alexandria, Mariout. Fayoum: ?. Lower Nile Valley and Delta: Kerdassa, Dokki, Subfamily EXORISTINAE Maadi. Sinai: El-Arish. Upper Nile Valley: Assiout. Tribe ERYCIINI Dates of collection: November to April. Genus AMELIBAEA Mesnil List of doubtful taxa (Nomina Dubia) described from Egypt: (see Herting and Dely-Draskovits 1993) – Amelibaea ?signifera (Villeneuve, 1929) – Catabucentes abdominalis Villeneuve, 1934b: 72. Genus APLOMYA Robineau-Desvoidy Egypt (Lower Egypt). – Eurigaster incisus Walker, 1871:340.Egypt (HotTamanib). – Aplomya confinis (Fallén, 1820) – Eurigaster leucomelas Walker, 1871: 339. Egypt. – Aplomya metallica (Wiedemann, 1824) – Frontina latifascia Walker, 1871: 340. Egypt (Wadi Genneh). Genus CADURCIELLA Villeneuve – Reaumuria olivieri Robineau-Desvoidy, 1830:82.Egypt. – Reaumuria virescens Robineau-Desvoidy, 1863b: 741. – Cadurciella tritaeniata (Rondani, 1859) Egypt. Genus CESTONIA Rondani Conclusion As a final conclusion, this catalogue treats a total of 72 – Cestonia cineraria Rondani, 1861 tachinid species belonging to 42 genera, 15 tribes and 4 – Cestonia rutilans Villeneuve, 1929 subfamilies in Egypt. Synonymies, type localities, type depositories, world distributions by biogeographic Genus CESTONIONERVA Villeneuve realm(s) and country, Egyptian localities and dates of collection are provided for all Egyptian taxa. – Cestonionerva petiolata (Villeneuve, 1910) Appendix Genus DRINO Robineau-Desvoidy Checklist of the Tachinidae of Egypt Subgenus PALEXORISTA Townsend Subfamily DEXIINAE Tribe DEXIINI – Drino (Palexorista) gilva (Hartig, 1838) Genus TRIXICEPS Villeneuve – Drino (Palexorista) imberbis (Wiedemann, 1830) – Drino (Palexorista) inconspicua (Meigen, 1830) – Trixiceps magnipalpis (Bezzi, 1922) Subgenus ZYGOBOTHRIA Mik Tribe VORIINI Genus HYPOVORIA Villeneuve – Drino (Zygobothria) atropivora (Robineau-Desvoidy, 1830) – Hypovoria hilaris (Villeneuve, 1913) Genus THECOCARCELIA Townsend Genus NANOPLAGIA Villeneuve – Thecocarcelia acutangulata (Macquart, 1851) – Nanoplagia sinaica (Villeneuve, 1909) Tribe ETHILLINI Genus ATYLOMYIA Brauer Genus PERISCEPSIA Gistel Subgenus PERISCEPSIA Gistel – Atylomyia albifrons Villeneuve, 1911 – Periscepsia (Periscepsia) handlirschi (Brauer and von Tribe EXORISTINI Bergenstamm, 1891) Genus ALLOPROSOPAEA Villeneuve El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 15 of 19 – Alloprosopaea efflatouni Villeneuve, 1923 Genus NEALSOMYIA Mesnil Genus CHAETORIA Becker – Nealsomyia triseriella (Villeneuve, 1929) – Chaetoria stylata Becker, 1908 Genus PSEUDOGONIA Brauer and Bergenstamm Genus CHETOGENA Rondani – Pseudogonia fasciata (Wiedemann, 1819) – Pseudogonia rufifrons (Wiedemann, 1830) – Chetogena aegyptiaca (Villeneuve, 1923) – Chetogena cercosa (Kugler, 1980) Genus RAMONELLA Kugler – Chetogena sinaica (Villeneuve, 1909) – Ramonella mesnili (Kugler, 1980) Genus EXORISTA Meigen. Subgenus ADENIA Robineau-Desvoidy Tribe WINTHEMIINI Genus NEMORILLA Rondani – Exorista (Adenia) rustica (Fallén, 1810) – Nemorilla floralis (Fallén, 1810) Subgenus EXORISTA Meigen Subfamily PHASIINAE – Exorista (Exorista) fasciata (Fallén, 1820) Tribe CYLINDROMYIINI – Exorista (Exorista) larvarum (Linnaeus, 1758) Genus BESSERIA Robineau-Desvoidy Subgenus PODOTACHINA Brauer and Bergenstamm – Besseria anthophila (Loew, 1871) – Exorista (Podotachina) sorbillans (Wiedemann, 1830) Genus CYLINDROMYIA Meigen Subgenus PTILOTACHINA Brauer and Bergenstamm – Cylindromyia intermedia (Meigen, 1824) – Cylindromyia rufipes (Meigen, 1824) – Exorista (Ptilotachina) rutilans Mesnil, 1970 – Exorista (Ptilotachina) xanthaspis (Wiedemann, 1830) Tribe LEUCOSTOMATINI Genus DIONOMELIA Kugler Genus MACULOSALIA Mesnil – Dionomelia hennigi Kugler, 1978 – Maculosalia maculosa (Villeneuve, 1909) Genus LEUCOSTOMA Meigen Tribe GONIINI Genus BLEPHARELLA Macquart – Leucostoma engeddense Kugler, 1966 – Leucostoma obsidianum (Wiedemann, 1830) – Blepharella versatilis (Villeneuve, 1910) Tribe PHASIINI Genus CLEMELIS Robineau-Desvoidy Genus PHASIA Latreille – Clemelis apicalis Villeneuve, 1923 – Phasia pusilla Meigen, 1824 Genus GONIA Meigen Subfamily TACHININAE Tribe ERNESTIINI – Gonia atra Meigen, 1826 Genus EURITHIA Robineau-Desvoidy – Gonia bimaculata Wiedemann, 1819 – Gonia capitata (De Geer, 1776) – Eurithia castellana (Strobl, 1906) – Gonia ornata Meigen, 1826 Genus LINNAEMYA Robineau-Desvoidy Genus MENDELSSOHNIA Kugler – Linnaemya comta (Fallén, 1810) – Mendelssohnia sinaica Kugler, 1971 – Linnaemya latigena Kugler, 1977 El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 16 of 19 – Linnaemya petiolata Kugler, 1971 London, London, United Kingdom; MHNL: Musée d’Histoire Naturelle de Lille, Lille, France; MNHN: Muséum National d’Histoire Naturelle, Paris, France; – Linnaemya soror Zimin, 1954 MRSN: Museo Regionale di Scienze Naturali, Torino [Turin], Italy; MZF: Museo Zoologico “La Specola”, Firenze [Florence], Italy; MZLU: Museum of Zoology, Tribe GRAPHOGASTRINI Lund University, Lund, Sweden; NHMW: Naturhistorisches Museum Wien, Wien [Vienna], Austria; NHRS: Naturhistoriska riksmuseet [Swedish Museum of Genus PHYTOMYPTERA Rondani Natural History], Stockholm, Sweden; NMBA: Naturhistorisches Museum der Benediktiner-Abtei Admont, Admont, Austria; PPDD: Ministry of Agriculture – Phytomyptera lacteipennis Villeneuve, 1934 Collection, Plant Protection Research Institute, Dokki, Giza, Egypt; SMF: Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany; SMNS: Staatliches Museum für Naturkunde, Stuttgart, Germany; Tribe MINTHOINI TAU: Tel Aviv University, Tel Aviv, Israel; USNM: National Museum of Natural History [formerly United States National Museum], Smithsonian Institution, Genus MINTHO Robineau-Desvoidy Washington, District of Columbia, USA; ZIN: Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia; ZMHB: Museum für Naturkunde – Mintho compressa (Fabricius, 1787) der Humboldt-Universität zu Berlin, Berlin, Germany; ZMUC: Zoological Museum, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark; ZMUH: Universität von Hamburg, Zoologisches Genus PLESINA Meigen Institut und Zoologisches Museum, Hamburg, Germany Biogeographic Realm abbreviations used throughout the catalogue – Plesina claripennis Mesnil, 1953 AF: Afrotropical; AU: Australasian; NE: Nearctic; NT: Neotropical; OR: Oriental; – Plesina deserticola Kugler, 1978 PA: Palaearctic Acknowledgements Genus ROSSIMYIOPS Mesnil I would like to thank Dr. Amnon Freidberg and Dr. Tatyana Novoselsky, Tel Aviv University, Israel for supplying me with some relevant papers. I am grateful to Dr. Ashraf Al-Torky, Plant Protection Institute, Ministry of Agricul- – Rossimyiops achilleae (Kugler, 1972) ture, Egypt, for his help throughout this study. – Rossimyiops magnificus (Kugler, 1972) Funding No funding. Tribe SIPHONINI Genus ACTIA Robineau-Desvoidy Availability of data and materials Data supporting the conclusions of this article are presented in the main manuscript. – Actia crassicornis (Meigen, 1824) Author’s contributions This is a single author study. The author read and approved the final Genus PERIBAEA Robineau-Desvoidy manuscript. Ethics approval and consent to participate – Peribaea orbata (Wiedemann, 1830) Not applicable. – Peribaea palaestina (Villeneuve, 1934) Consent for publication Not applicable. Genus SIPHONA Meigen Subgenus SIPHONA Meigen, 1803 Competing interests The author declares that he has no competing interests. – Siphona (Siphona) efflatouni Mesnil, 1960 Publisher’sNote Springer Nature remains neutral with regard to jurisdictional claims in Tribe TACHININI. published maps and institutional affiliations. Genus PELETERIA Robineau-Desvoidy. Received: 18 December 2017 Accepted: 19 March 2018 Peleteria meridionalis (Robineau-Desvoidy, 1830). Abbreviations References List of abbreviations used throughout the catalogue Becker T (1908) Dipteren der Kanarischen Inseln. Mitt Zool Mus Berlin 4:1–180 C.: Central; E.: East; Eur.: Europe; ICZN: International Commission on Zoological Becker T (1910) Dipteren aus Südarabien und von der Insel Sokótra. Kaiserl Akad Nomenclature; I., Is.: Island, islands; M.: Middle; N.: North; Ne.: Northeast; Wiss Wien, Math Naturwiss Kl, Denkschr 71:131–160 Nw.: Northwest; nr.: near; Pal.: Palaearctic; S.: South; St.: Saint; var.: Variety; Bezzi M (1922) Contributo allo studio della fauna Libica. Ditteri di Cirenaica W.: West raccolti dal Rev Miss Don Vito Zanon Mem Soc entomol ital 1: 140–157 Bezzi M, Lamb CG (1926) Diptera (excluding Nematocera) from the island of Museum abbreviations used throughout the catalogue Rodriguez. Trans Entomol Soc Lond 58(1925):537–573 BMNH: Natural History Museum [formerly British Museum (Natural History)], Bigot JMF (1881) Diptères nouveaux ou peu connus. 17e partie. XXVI. 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Trudy Vses Entomol Obshch 48:230–334 http://www.deepdyve.com/assets/images/DeepDyve-Logo-lg.png Egyptian Journal of Biological Pest Control Springer Journals

Catalogue of the Tachinidae of Egypt (Diptera: Oestroidea)

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Abstract

Tachinid flies are an important group of parasitoids in their larval stage, and all their hosts are of the Arthropoda, almost exclusively other insects, including important insect pests in agriculture and forestry. All known Egyptian taxa of the family Tachinidae are systematically catalogued. Synonymies, type localities, type depositories, world distributions by biogeographic realm(s) and country, Egyptian localities, and dates of collection are provided. A total of 72 tachinid species belonging to 42 genera, 15 tribes, and 4 subfamilies has been treated. Keywords: Tachinid flies, Egyptian taxa, World distribution, Egyptian localities, Dates of collection Background agriculture and forestry. They typically parasitize phytopha- Tachinidae are a large and cosmopolitan family of flies gous larvae of Lepidoptera and Coleoptera or nymphs of within the superfamily Oestroidea. It is the second largest Hemiptera and Orthoptera. Consequently, tachinid flies family in the order Diptera (Irwin et al. 2003), with some have been successfully applied in programs of biological 1500 recognized genera (O’Hara 2016) and more than control against different insect pests (Stireman et al. 2006; 8500 described species (O’Hara 2013) worldwide. How- O’Hara 2008 and Cerretti and Tschorsnig 2010). ever, the estimated true diversity of the family is probably No comprehensive taxonomic studies on the family double the number of the currently known species, mak- Tachinidae have been carried out in Egypt before. Only one ing this family a good candidate for the first largest family published study, Steyskal and El-Bialy (1967), listed the of Diptera (O’Hara, 2008 and 2013 and Pape et al. 2011). Egyptian species of dipterous families including Tachinidae Tachinid flies range in size from 2 to 20 mm, and most of based on literature and specimens deposited in the main them are larger than a house fly and noticeably more bris- Egyptian insect collections. However, despite its import- tly, especially on thoracic dorsum and third to fifth abdom- ance, the list included only the names of species within the inal segments; however, across the family there is a family ordered alphabetically without referring to their tremendous variety of shapes, colors, and degree of brist- suprageneric classification or to other important data such ling. In addition to features shared with other families of as original descriptions, type localities, synonymies, world the super family Oestroidea (Rhinophoridae, Sarcophagi- distribution, type depositories, and local distribution. An- dae, Calliphoridae, and Oestridae), adult tachinids can be otherstudy,anunpublished Ph.D.thesisbyDr. Fathy recognized by the developed postscutellum (a rounded Negm (Plant Protection Institute, Ministry of Agriculture) bulge below the hind part of the thoracic dorsum) (O’Hara was completed in 1987 and reviewed the taxonomy of 2008). The family is divided into four subfamilies: Dexiinae, Egyptian Tachinidae. The author mainly adopted the classi- Exoristinae, Phasiinae, and Tachininae. All four subfamilies fication of the Palaearctic catalogue of Herting (1984). are represented in Egypt as shown in the present results. All missing data and deficiencies in previous Egyptian Tachinid flies are nectar or honeydew feeders as adults studies have been perfected with an updated taxonomy and are effective pollinators (Tooker et al. 2006). However, of the Egyptian Tachinidae in the present catalogue. they are an important group of parasitoids in their larval The present study is one in a series of planned studies stage, and all their hosts are of the phylum Arthropoda, al- on different Egyptian dipteran families aiming to cata- most exclusively other insects, including insect pests in logue the order Diptera as a whole in Egypt. Egypt, as a part of the Great Desert Belt, is characterized Correspondence: elhawagry@gmail.com by a warm and almost rainless climate (El-Hawagry 2017). Entomology Department, Faculty of Science, Cairo University, Giza, Egypt © The Author(s). 2018 Open Access This article is distributed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted use, distribution, and reproduction in any medium, provided you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 2 of 19 Ecologists divide Egypt into eight ecological zones: Coastal and written in bold uppercase letters. Species-group Strip, Lower Nile Valley and Delta, Upper Nile Valley, headings are left-justified and written in bold italicized Fayoum, Eastern Desert, Western Desert, Sinai, and Gebel lowercase letters except the first letter which is written Elba (Fig. 1). These zones have a greater affiliation to the uppercase. Authorship of genera, subgenera, and species Palaearctic Region except for the Gebel Elba, the southeast- are written in lowercase letters except the first letter ern triangle of Egypt, which has a greater affiliation to the which is written in uppercase. Taxonomically valid Afrotropical Region (El-Hawagry and Gilbert 2014). genus-group names (senior synonyms) are listed again in bold italicized lowercase letters (except the first upper- Material and methods case letter) and left-justified under the headings followed The present catalogue is based mainly on specimens col- by the reference including author, year, and journal lected by the author and his co-workers from different pages. Taxonomically valid species-group names com- Egyptian localities, in addition to data compiled from bined with their original genera (senior synonyms) are specimens preserved in the main Egyptian insect collec- listed again in regular italicized lower case letters (except tions, especially EFC and PPDD. A great deal of infor- the first uppercase letter) and left-justified under the mation, including synonymies and distributional data is headings followed by the reference including author, obtained from relevant literature and website databases year, and journal pages. Type species for all nomenclatu- as well. Of the most important literature reviewed are rally available genus-group names are given after the ref- Kugler (1980a); Herting (1984); Cerretti and Freidberg erence line, followed by method of their fixation. For 2009;O’Hara et al. (2009); O’Hara and Cerretti (2016); each genus and species-group name, associated syno- and Pape and Thompson (2017). nyms are listed in a chronological order. They are writ- ten in regular italicized lowercase letters (except the first Scope uppercase letter), followed by the reference and other This catalogue treats all names of taxa of the family data as in senior taxa. The rank of types that currently Tachinidae recorded from Egypt. exist (e.g., syntype, holotype, lectotype) is given after the reference of each species-group taxon, followed by its Arrangement of taxa location (depository) in museums, which is given as All taxa are arranged alphabetically. Old world syno- abbreviations in parenthesis. The type locality of each nyms, including all available and unavailable names, are species-group taxon is given at the end after its type chronologically listed. depository. Countries and islands of type localities are usually broken down to states, provinces, archipelagos, Typographical treatment of names ecological zones, towns, and villages. These “sublocal- Family-group headings are centered and written in bold ities” are placed in parentheses after the main locality, e. uppercase letters. Genus-group headings are left-justified g., “Egypt (Sinai)” . The sublocalities may be more broken down to “smaller sublocalities”. These “smaller sublocalities” are written after a colon following the sub- locality, e.g., “Egypt (Sinai: Tor)”. Distribution A complete world distribution by biogeographic realm(s) and country of each tachinid species, as is known from the litera- ture, is listed in alphabetical order. Distribution localities pre- ceded by a question mark (?) are considered questionable. Egyptian localities and dates of collection The Egyptian localities from which each tachinid species is collected and dates of collection are given to map a picture for the distribution and activity periods of the species in the different ecological zones of Egypt. Local- ities within each ecological zone are arranged alphabetic- ally and written after a colon following the ecological zone, e.g., “Coastal Strip: Mariout”. Spelling of locality Fig. 1 Map of Egypt showing the ecological zones (taken from El- names in Egypt almost follows El-Hawagry and Gilbert Hawagry and Gilbert 2014) (2014) and El-Hawagry (2015 and 2017). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 3 of 19 The catalogue Plagia hilfii sinaica Villeneuve in Hermann and This catalogue presents four tachinid subfamilies as rep- Villeneuve, 1909: 157. Holotype female (CNC). Egypt resented in Egypt, and the number of taxa treated in the (Sinai). catalogue can be summarized in table. Distribution: PA: Algeria, China, Egypt, S. Europe (Albania, Bosnia and Herzegovina, Bulgaria, Croatia, France, Greece, Italy, Malta, Monaco, Montenegro, Subfamily Tribes Genera Species Macedonia, San Marino, Serbia, Slovenia), Israel, Dexiinae 2 6 6 Kazakhstan,Morocco,Russia (W.Russia,E.Siberia), Exoristinae 5 21 37 Transcaucasia. Phasiinae 3 5 7 Egyptian localities: Sinai:? Tachininae 5 10 16 Dates of collection: April. Doubtful taxa 6 Genus PERISCEPSIA Gistel Total number 15 42 72 Scopolia Robineau-Desvoidy, 1830: 268 (junior homo- nym of Scopolia Hübner). Type species: Musca carbo- naria Panzer, 1798, by subsequent designation of Zetterstedt, 1844: 1239). Periscepsia Gistel, 1848:x(nomen novum for Scopolia Subfamily DEXIINAE Robineau-Desvoidy, 1830). Tribe DEXIINI Phoricheta Rondani, 1861a:8(nomen novum for Genus TRIXICEPS Villeneuve Scopolia Robineau-Desvoidy, 1830). Trixiceps Villeneuve, 1936: 329. Type species: Ramonda Robineau-Desvoidy, 1863a: 790. Type spe- Trixiceps aegyptiaca Villeneuve, 1936, by monotypy. cies: Ramonda fasciata Robineau-Desvoidy, 1863 Trixiceps magnipalpis (Bezzi, 1922) (= Tachina spathulata Fallén, 1820) by original Paraprosena magnipalpis Bezzi, 1922: 154. Type(s) not designation. located. Libya (Benghazi). Subgenus PERISCEPSIA Gistel —Nominotypical Trixiceps aegyptiaca Villeneuve, 1936: 329. Egypt subgenus (Ikingi Mariout). Periscepsia (Periscepsia) handlirschi (Brauer and Distribution: PA: Egypt, Europe, Libya. von Bergenstamm, 1891) Egyptian localities: Coastal Strip: Mariout, Mersa Phorichaeta handlirschii Brauer and Bergenstamm, Matruh. 1891: 52. Unspecified type(s) (NHMW). Italy (Trentino: Dates of collection: April to August. Trafoi). Note: The specific epithet was spelled handlirschii in Tribe VORIINI the original description but was subsequently changed to Genus HYPOVORIA Villeneuve handlirschi (O’Hara 2009). Hypovoria Villeneuve, 1913a: 510 (as subgenus of Distribution: PA: China, Egypt, Israel, Europe. Voria Robineau-Desvoidy 1830). Type species: Voria Egyptian localities: Coastal Strip: Burg El-Arab, Mariout. (Hypovoria) hilaris Villeneuve, 1913, by monotypy. Dates of collection: February to October. Hypovoria hilaris (Villeneuve, 1913) Genus UCLESIA Girschner Voria (Hypovoria) hilaris Villeneuve, 1913a: 510. Uclesia Girschner, 1901: 69. Type species: Uclesia Holotype female (CNC). Tunisia (Sfax). fumipennis Girschner, 1901, by monotypy. Distribution: PA: Canary Is., C. Asia (Kyrgyzstan, Uclesia petiolata (Villeneuve, 1929) Tajikistan, Turkmenistan, Uzbekistan), China, Egypt, Nanoplagia petiolata Villeneuve, 1929a: 45. Holotype S. Europe (Albania, Bosnia and Herzegovina, Bulgaria, female (?EFC, not located). Egypt (Wadi Rishrash). Croatia, France, Greece, Italy, Malta, Monaco, Distribution: PA: Algeria, Tunisia, Egypt, Russia, Montenegro, Macedonia, San Marino, Serbia, Tadzhikistan. Slovenia), W. Europe (Andorra, Portugal (including Egyptian localities: Eastern Desert: Wadi Rishrash. Azores, excluding Madeira), Spain), Israel, Mongolia, Dates of collection: March. Russia (E. Siberia), Tunisia, Transcaucasia. Genus WAGNERIA Robineau-Desvoidy Egyptian localities: Coastal Strip: El-Marg, Mariout. Wagneria Robineau-Desvoidy, 1830: 126. Type spe- Dates of collection: February to July. cies: Wagneria gagatea Robineau-Desvoidy, 1830,by Genus NANOPLAGIA Villeneuve monotypy. Nanoplagia Villeneuve, 1929a: 45. Type species: Wagneria cunctans (Meigen, 1824) Plagia hilfii Strobl, 1902, by original designation. Tachina cunctans Meigen, 1824: 419. Holotype male Nanoplagia sinaica (Villeneuve, 1909) (MNHN). Germany. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 4 of 19 Wagneria riedeli Villeneuve, 1937: 405. Germany Exorista aenescens Macquart, 1850: 409. France (Frankfurt). (Lestrem). Distribution: PA: Egypt, Europe, Israel. Exorista longicornis Perris, 1852: 206. France (Grandes Egyptian localities: Coastal Strip: El-Burg, Mariout, Landes). Nuzha. Eastern Desert: Suez Road. Lower Nile Valley Tachina bizonata Zetterstedt, 1859: 6119. Sweden and Delta: Dokki. Sinai: Ein Mouussa. (Skäne: Sjobo). Dates of collection: January to July. Distribution: AF: ?Malawi, Yemen. OR: Orien. China [see O’Hara and Cerretti 2016: 19]. PA: Canary Is., C. Asia (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), Subfamily EXORISTINAE Egypt, Europe, Israel, Japan, Korea, Kuril Is., Madeira, Tribe ERYCIINI Mongolia, Pal. China [see O’Hara and Cerretti 2016: 19], Genus AMELIBAEA Mesnil Russia, Transcaucasia. Amelibaea Mesnil, 1955: 454 (as subgenus of Egyptian localities: Lower Nile Valley and Delta: Abu Phebellia Robineau-Desvoidy, 1846). Type species: Par- Rawash, Koubba. exorista tultschensis Brauer and Bergenstamm, 1891,by Dates of collection: February to April. monotypy. Aplomya metallica (Wiedemann, 1824) Amelibaea ?signifera (Villeneuve, 1929) Tachina metallica Wiedemann, 1824: 46. Lectotype Exorista signifera Villeneuve, 1929a: 44. Holotype male male (ZMUC). India. [by fixation of Townsend (1933: (?EFC, not located). Egypt (Mariout). 470)]. Remarks: The identity of this species is not clear since Tachina nigriventris Wiedemann, 1824: 43. India. the male holotype is apparently lost (Cerretti and Tachina notata Wiedemann, 1830: 653. Sudan (Nubia Freidberg 2009). Region [as “Nubien”]). Distribution: PA: Egypt, Israel (Cerretti and Freidberg Tachina socia Wiedemann, 1830: 654. Sudan (Nubia 2009). Region [as “Nubien”]). Egyptian localities: Coastal Strip: Mariout. Phorocera eucalypta Loew, 1852: 659. Mozambique Dates of collection: January. (Tete [as “Tette”]). Genus APLOMYA Robineau-Desvoidy Parexorista laeviventris Wulp, 1893: 173. Indonesia Aplomya Robineau-Desvoidy, 1830: 184. Type species: (Jawa). Aplomya zonata Robineau-Desvoidy, 1830 (= Tachina Tricholyga impexa Villeneuve, 1916: 494. D.R. Congo confinis Fallén, 1820), by subsequent designation of and South Africa (Eastern Cape). Robineau-Desvoidy (1863a: 459, 460). Distribution: AF: Widespread. OR: Widespread. Prozenillia Villeneuve, 1916: 487. Type species: Palaearctic: Egypt, Israel, Japan, Pal. China [see O’Hara Prozenillia distans Villeneuve, 1916, by monotypy. and Cerretti 2016: 19]. Wiedemanniomyia Townsend, 1933: 469. Type Egyptian localities: Upper Nile Valley: Asswan. species: Tachina metallica Wiedemann, 1824,by Dates of collection: Unknown. original designation. Genus CADURCIELLA Villeneuve Aplomyiella Mesnil, 1939: 31. Type species: Tricholyga Cadurciella Villeneuve, 1927: 120. Type species: impexa Villeneuve, 1916 (= Tachina metallica Cadurciella rufipalpis Villeneuve, 1927, by monotypy. Wiedemann, 1824), by original designation. Cadurciella tritaeniata (Rondani, 1859) Atricholyga Villeneuve, 1939: 9. Type species: Exorista tritaeniata Rondani, 1859: 147. Unspecified Tricholyga impexa Villeneuve, 1916 (= Tachina type(?MZF). Italy (Parma). metallica Wiedemann, 1824), by monotypy. Distribution: PA: Egypt, Europe (northwards to Aplomya confinis (Fallén, 1820) Scotland, S. Norway, Sweden, Japan. Tachina confinis Fallén, 1820b: 32. Syntypes, males Egyptian localities and dates of collection are not precisely and females (NHRS and/or MZLU). Sweden (Gotland). known. Recorded by Steyskal and El-Bialy (1967: 48). Phryxe zonata Robineau-Desvoidy, 1830: 159. France. Genus CESTONIA Rondani Phryxe servillei Robineau-Desvoidy, 1830: 159. France Cestonia Rondani, 1861a: 105. Type species: Cestonia (Paris). cineraria Rondani, 1861, by monotypy. Aplomya nitens Robineau-Desvoidy, 1830: 185. France Cestonia cineraria Rondani, 1861 (Yonne: St. Sauveur). Cestonia cineraria Rondani, 1861a: 106. Syntypes, Aplomya zonata Robineau-Desvoidy, 1830: 185. France males and females (MZF). Italy (Parma). (Paris). Distribution: PA: Egypt, Europe (Mediterranean, Tachina bicingulata Zetterstedt, 1844: 1133. Sweden northwards to S. France, Switzerland, and Italy), (Skäne: Esperod). Israel. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 5 of 19 Egyptian localities: Coastal Strip: Nuzha. Eastern Egyptian localities and dates of collection are not Desert: Wadi Hoff. Lower Nile Valley and Delta: Ezbet precisely known. El-Nakhl. Sinai: Wadi El-Tih. Drino (Palexorista) inconspicua (Meigen, 1830) Dates of collection: March to July. Tachina inconspicua Meigen, 1830: 369. Unspecified Cestonia rutilans Villeneuve, 1929 types, females (MNHN). Germany (Berlin). Cestonia rutilans Villeneuve, 1929b: 102. Syntypes, 1 Tachina bimaculata Hartig, 1838: 286. Germany. male and 1 female, not located. Egypt (Cairo). Tachina flavoscutellata Zetterstedt, 1844: 1042. Sweden. Distribution: AF: Senegal, Yemen. PA: Egypt, Libya. Masicera cursitans Rondani, 1861a: 21. Italy (Parma). Egyptian localities and dates of collection are not Sturmia lophyri Robineau-Desvoidy, 1863b: 892. precisely known. Germany. Genus CESTONIONERVA Villeneuve Drino inusta Mesnil, 1949: 25. Poland. Cestonionerva Villeneuve, 1929a: 43. Type species: Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Conogaster petiolata Villeneuve, 1910, by subsequent Turkmenistan, Uzbekistan), China, Egypt, Europe, designation of Townsend (1936: 137). Russia (W. Russia, W. Siberia), Taiwan, Transcaucasia. Cestonionerva petiolata (Villeneuve, 1910) Egyptian localities and dates of collection are not Conogaster petiolata Villeneuve in Becker, 1910: 144. precisely known. Holotype female (NHMW, not located). Yemen (Socotra). Subgenus ZYGOBOTHRIA Mik Cestonionerva proxima Villeneuve, 1929a: 43. Egypt Zygobothria Mik, 1891: 193. Type species: Sturmia (Dahshour). atropivora Robineau-Desvoidy, 1830, by original Distribution: AF: U.A. Emirates, Yemen. PA: Canary designation. Is., C. Asia (Kyrgyzstan, Tajikistan, Turkmenistan, Formosodoria Townsend, 1933: 475. Type species: Uzbekistan), Israel, Mongolia, North Africa, Pal. China Sturmia (Argyrophylax) dilabida Villeneuve, 1916 (= [see O’Hara and Cerretti 2016: 19]. Meigenia ciliata van der Wulp, 1881), by original Egyptian localities: Lower Nile Valley and Delta: Dahshour. designation. Dates of collection: April. Drino (Zygobothria) atropivora (Robineau-Desvoidy, Genus DRINO Robineau-Desvoidy 1830) Drino Robineau-Desvoidy, 1863a: 250. Type species: Sturmia atropivora Robineau-Desvoidy, 1830: 171. Drino volucris Robineau-Desvoidy, 1863 (= Tachina lota Syntypes, males and females (MNHN). France. Meigen, 1824), by original designation. Sturmia masakensis Curran, 1927c: 117. Uganda Subgenus PALEXORISTA Townsend (Masaka). Palexorista Townsend, 1921: 134. Type species: Distribution: AF: Widespread. AU: Australia. OR: Tachina succini Giebel, 1862 (as “Tichina succini India, Indonesia, Laos, Malaysia, Orien. China [see Giebel”), by monotypy. O’Hara and Cerretti, 2016: 19], Ryukyu Is., Sri Lanka. Prosturmina Mesnil, 1970b: 110 (as subgenus of Drino PA: Algeria,CanaryIs.,C.Asia(Kyrgyzstan, Robineau-Desvoidy). Type species: Sturmia vigilans Tajikistan, Turkmenistan, Uzbekistan), Egypt, all Villeneuve, 1933 (= Sturmia pulchra Curran, 1927), Europe (exceptBritishIs.,Scand.),Israel,Japan,Pal. by original designation. China[seeO’Hara and Cerretti 2016: 19], Morocco, Drino (Palexorista) gilva (Hartig, 1838) Russia, Transcaucasia, Tunisia. Tachina gilva Hartig, 1838: 288. Syntypes, males and Egyptian localities: Coastal Strip: Ramleh. Eastern females, not located. Germany. Desert: Ismailia, Suez Road. Lower Nile Valley and Drino aurora Mesnil, 1949: 15. Japan. Delta: El-Qubba, Dekernes, Maadi, Shubra. Distribution: PA: Egypt, Europe, Japan, Transcaucasia. Dates of collection: April to July. Egyptian localities and dates of collection are not Genus THECOCARCELIA Townsend precisely known. Recorded from Egypt by Steyskal and Thecocarcelia Townsend, 1933: 471. Type species: El-Bialy (1967: 48). Argyrophylax pelmatoprocta Brauer and Bergenstamm, Drino (Palexorista) imberbis (Wiedemann, 1830) 1891 (= Masicera acutangulata Macquart, 1851), by Tachina imberbis Wiedemann, 1830: 317. Syntypes, original designation. males (ZMUH, lost). Egypt. Thelycarcelia Townsend, 1933: 475. Type species: Sturmia zonata Curran, 1927b: 336. Uganda (Entebbe). Thelycarcelia thrix Townsend, 1933 (= Sturmia Distribution: AF: D.R. Congo, Kenya, Malawi, South sumatrana Baranov, 1932), by original designation. Africa, Sudan, Tanzania, U.A. Emirates, Uganda, Yemen. Thecocarcelia acutangulata (Macquart, 1851) PA: Canary Is., C. Asia (Kyrgyzstan, Tajikistan, Masicera acutangulata Macquart, 1851a: 478. Turkmenistan, Uzbekistan), Egypt, Iran, Israel, North Unspecified type, female (?MHNL). Switzerland (Chur Africa, Turkey, Turkmenia. [as “Coire”]). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 6 of 19 Masicera incedens Rondani, 1861a: 22. Italy (Parma). Robineau-Desvoidy, 1830 (= Tachina obliquata Fallén 1810), Argyrophylax pelmatoprocta Brauer and Bergenstamm, by subsequent designation of Robineau-Desvoidy (1863a:553). 1891: 344. Austria (Bisamberg: nr. Vienna). Chetogena Rondani, 1856: 68. Type species: Salia Distribution: AF: Widespread. PA: Palaearctic: Egypt, rondaniana Villeneuve, 1931. all Europe (except Turkey), Japan, Transcaucasia. Spoggosia Rondani, 1859: 182. Type species: Spoggosia Egyptian localities and dates of collection are not occlusa Rondani, 1859 (=Tachina obliquata Fallén 1810), precisely known. by monotypy. Stomatomyia Brauer and Bergenstamm, 1889: 98. Tribe ETHILLINI Type species: Chetogena filipalpis Rondani, 1859,by Genus ATYLOMYIA Brauer subsequent designation of Brauer (1893: 483). Atylomyia Brauer, 1898: 525. Type species: Atylomyia Chetogena aegyptiaca (Villeneuve, 1923) loewii Brauer, 1898, by monotypy. Tachinosalia aegyptiaca Villeneuve, 1923: 91. Holotype Atylomyia albifrons Villeneuve, 1911 male (EFC). Egypt. Atylomyia albifrons Villeneuve, 1911: 86. Lectotype Distribution: PA: Egypt, Israel. female (CNC). Egypt (Helwan). Egyptian localities: Coastal Strip: Mariout. Atylomyia rungsi Mesnil, 1962: 778. Holotype. Dates of collection: March. Morocco (Ait Melloul). Chetogena cercosa (Kugler, 1980) Distribution: PA: China, Egypt, Israel, W. Europe Chaetogena cercosa Kugler, 1980a: 31. Holotype male (Andorra, Portugal (including Azores, excluding (TAU). Israel (Elat). Madeira), Spain). Distribution: AF: U. A. Emirates. PA: Egypt, Israel. Egyptian localities: Lower Nile Valley and Delta: Egyptian localities: Sinai: Serabit El-Khadim. Helwan. Dates of collection: April. Dates of collection: January and February. Chetogena sinaica (Villeneuve, 1909) Stomatomyia acuminata var. sinaica Villeneuve in Tribe EXORISTINI Hermann and Villeneuve, 1909: 157. Holotype female Genus ALLOPROSOPAEA Villeneuve (CNC). Egypt (Sinai). Alloprosopaea Villeneuve, 1923: 89. Type species: Distribution: PA: Egypt, Israel. Alloprosopaea efflatouni Villeneuve, 1923, by monotypy. Egyptian localities: Sinai: South Sinai. Alloprosopaea efflatouni Villeneuve, 1923 Dates of collection: ?April. Alloprosopaea efflatouni Villeneuve, 1923: 90. Genus EXORISTA Meigen Unspecified type(s) (CNC). Egypt (Wadi Hoff). Exorista Meigen, 1803: 280. Type species: Musca Distribution: PA: Egypt, Israel. larvarum Linnaeus, 1758, by monotypy. Egyptian localities: Eastern Desert: Wadi Hoff. Lower Subgenus ADENIA Robineau-Desvoidy Nile Valley and Delta: Dahshour. Adenia Robineau-Desvoidy, 1863a: 1041. Type species: Dates of collection: April. Tachina grisea Robineau-Desvoidy, 1830 (= Tachina rustica Genus CHAETORIA Becker Fallén, 1810), by original designation. Chaetoria Becker, 1908: 113. Type species: Chaetoria Staegeria Robineau-Desvoidy, 1863a: 972 (junior homo- stylata Becker, 1908, by monotypy. nym of Staegeria Rondani 1856). Type species: Tachina Clistorrhinia Bezzi in Bezzi and Lamb, 1926: 570. pratensis Robineau-Desvoidy, 1830,byoriginaldesignation. Type species: Clistorrhinia aurifrons Bezzi, 1926,by Chaetotachina Brauer and Bergenstamm, 1889: 98. monotypy. Type species: Tachina rustica Fallén, 1810, by monotypy. Chaetoria stylata Becker, 1908 Exorista (Adenia) rustica (Fallén, 1810) Chaetoria stylata Becker, 1908: 114. Lectotype female Tachina rustica Fallén, 1810: 264. Lectotype male (ZMHB). Canary Islands (Tenerife). (NHRS). Sweden (Skåne). Distribution: AF: Botswana, Mozambique, Nigeria, Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Senegal, U.A. Emirates, Yemen. PA: Algeria, Canary Is., C. Turkmenistan, Uzbekistan), China, Egypt, all Europe, M. Asia (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), East, Mongolia, Russia, Taiwan, Transcaucasia. Egypt, Europe, Morocco, Tunisia. Egyptian localities: Eastern Desert: Wadi Hoff. Egyptian localities: Coastal Strip: Ramleh. Upper Nile Dates of collection: March to June. Valley: Luxur. Subgenus EXORISTA Meigen —Nominotypical Dates of collection: January to September. subgenus Genus CHETOGENA Rondani Exorista Meigen, 1803: 280. Type species: Musca Salia Robineau-Desvoidy, 1830: 108 (junior homonym larvarum Linnaeus, 1758 (as “Musca larvarum Fabr.”), of Salia Hübner, 1818). Type species: Salia echinura by monotypy. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 7 of 19 Exorista (Exorista) fasciata (Fallén, 1820) Distribution: PA: Egypt, Israel. Tachina fasciata Fallén, 1820a: 5. Syntypes, males and Egyptian localities: Sinai. females (NHRS and/or MZLU). Sweden (Skåne, Äsperöd Dates of collection: April. [as “Esperöd”] and Abusa). Exorista (Ptilotachina) xanthaspis (Wiedemann, 1830) Chetogena segregata Rondani, 1859: 181. Italy (Parma Tachina xanthaspis Wiedemann, 1830: 314. Syntypes, and Sicily); Malta. males and females (SMF). Sudan (Nubia Region [as Distribution: NE: Yukon, introduced and established in “Nubien”]). Ontario, Québec and New England. OR: India. PA: Tachina pyrrhocera Wiedemann, 1830: 314. Sudan China, Egypt, Europe, Japan, M. East, Mongolia, Russia, (Nubia Region [as “Nubien”]). Taiwan, Transcaucasia. Tachina fallax pseudofallax Villeneuve, 1920: 151. Egyptian localities: Upper Nile Valley: Assiout. South Africa (Eastern Cape). Dates of collection: March. Larvaevora (Ptilotachina) fallax aethiopica Exorista (Exorista) larvarum (Linnaeus, 1758) Rohdendorf, 1931: 348. Sudan (Wad Medani). Musca larvarum Linnaeus, 1758: 596. Unspecified Distribution: AF: Madagascar, Seychelles, Sudan, U.A. Type(s) (LSUK). Europe. Emirates, Yemen. AU: N. Australasian. OR: India, Tachina moreti Robineau-Desvoidy, 1853: 534. France Indonesia, Orien. China [see O’Hara and Cerretti 2016: (Auxerre). 19], Ryukyu Is., Taiwan. PA: C. Asia (Kyrgyzstan, Distribution: NE: New England, Ontario, Québec, Tajikistan, Turkmenistan, Uzbekistan), Egypt, all Europe Yukon. OR: India. PA: C. Asia (Kyrgyzstan, Tajikistan, (except British Is., Scand.), Israel, Mongolia, Pal. China Turkmenistan, Uzbekistan), China, all Europe, Japan, M. [see O’Hara and Cerretti 2016: 19], Russia, East, Mongolia, North Africa, Russia, Taiwan, Transcaucasia. Transcaucasia. Egyptian localities: Lower Nile Valley and Delta: Abu Egyptian localities: Coastal Strip: Alexandria, Mariout, Rawash. Sinai: St. Katherine. Nuzha. Eastern Desert: Ismailia. Lower Nile Valley and Dates of collection: February to August. Upper Nile Delta: Bahtim, Dessouk, Dokki, Embaba, Sakha, Tanta. Valley: Asswan. Fayoum: ?. Sinai: El-Arish. Upper Nile Valley: Sohag. Genus MACULOSALIA Mesnil Dates of collection: Throughout the year. Maculosalia Mesnil, 1946: 62 (as subgenus of Subgenus PODOTACHINA Brauer and Bergenstamm Spoggosia Rondani 1859). Type species: Deuterammobia Podotachina Brauer and Bergenstamm, 1891: 350. maculosa Villeneuve, 1909, by monotypy. Type species: Tachina sorbillans Wiedemann, 1830,by Maculosalia maculosa (Villeneuve, 1909) subsequent designation of Townsend (1916: 8). Deuterammobia maculosa Villeneuve, 1909: 80. Exorista (Podotachina) sorbillans (Wiedemann, 1830) ?Holotype (?MNHN). Egypt (Sinai). Tachina sorbillans Wiedemann, 1830: 311. Lectotype Distribution: PA: Egypt, Iran, Tunisia. male (NHMW). Canary Islands (Tenerife). Egyptian localities: Sinai: ? Distribution: AF: Cameroon, Canary Is., D.R. Congo, Dates of collection: Unknown. Kenya, Malawi, Nigeria, Sierra Leone, Uganda. OR: India, Indonesia, Nepal, Orien. China [see O’Hara and Tribe GONIINI Cerretti 2016: 19], Philippines, Ryukyu Is., Sri Lanka, Genus BLEPHARELLA Macquart Taiwan, Thailand, Vietnam. Australasian: Australia, N. Blepharella Macquart, 1851b: 176. Type species: Australasian. PA: C. Asia (Kyrgyzstan, Tajikistan, Blepharella lateralis Macquart, 1851, by original Turkmenistan, Uzbekistan), Egypt, Europe, Israel, Japan, designation. Mongolia, China. Blepharella versatilis (Villeneuve, 1910) Egyptian localities and dates of collection are not Sturmia versatilis Villeneuve, 1910a: 253. Unspecified precisely known. Recorded from Egypt by Steyskal and type(s), 1 male (CNC). D.R. Congo. El-Bialy (1967: 49). Distribution: AF: D.R. Congo, Malawi, Nigeria, Sudan. Subgenus PTILOTACHINA Brauer and PA: ?Egypt. Bergenstamm Note: Villeneuve (1913b: 29) recorded this species Ptilotachina Brauer and Bergenstamm, 1891:350. from Egypt, based on a male in BMNH, however this Type species: Exorista florentina Herting, 1975, by record needs confirmation (O’Hara and Cerretti 2016). fixation of O’Hara et al.(2009: 94) under Article 70.3.2 Genus CLEMELIS Robineau-Desvoidy of the Code (ICZN, 1999). Clemelis Robineau-Desvoidy, 1863a: 481. Type spe- Exorista (Ptilotachina) rutilans Mesnil, 1970 cies: Zenillia ciligera Robineau-Desvoidy, 1830 Exorista rutilans Mesnil, 1970a: 113. Holotype male (= Tachina pullata Meigen, 1824), by original (MNHN). Egypt (Sinai). designation. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 8 of 19 Clemelis apicalis Villeneuve, 1923 Kom Osheem. Lower Nile Valley and Delta: Abu Clemelis apicalis Villeneuve, 1923: 92. Holotype male Rawash, Borkash, El-Marg, Ezbet El-Nakhl, Kafr (?EFC, not located). Egypt [as “Lower Egypt”]. Hakim, Pyramids, Shubra, Tanta, Zaqaziq. Distribution: PA: Egypt. Dates of collection: February to September. Egyptian localities: Coastal Strip: Mariout. Gonia capitata (De Geer, 1776) Dates of collection: April. Musca capitata De Geer, 1776: 23. Syntypes, Genus GONIA Meigen unspecified number and sex (?NHRS). Sweden. Salmacia Meigen, 1800: 38. Name suppressed by Rhedia testacea Robineau-Desvoidy, 1830: 75. France ICZN (1963: 339). (Paris). Gonia Meigen, 1803: 280. Type species: Gonia Gonia trifaria Zeller, 1842: 841. Austria (Wien), bimaculata Wiedemann, 1819, by subsequent Poland (Reinerz and Szczecin). designation of Sabrosky and Arnaud Jr (1965: 1075). Gonia maritima Perris, 1847: 495. France. Reaumuria Robineau-Desvoidy, 1830: 79. Type spe- Distribution: PA: China, Egypt, Europe, Mongolia, cies: Musca capitata De Geer, 1776, by subsequent des- Russia (W. Russia, W. Siberia), Transcaucasia. ignation of Robineau-Desvoidy (1863a: 733). Egyptian localities: Common throughout the country, Pissemya Robineau-Desvoidy, 1851b: 318. Type spe- particularly in: Coastal Strip: El-Burg, Mariout. Fayoum: ?. East- cies: Gonia atra Meigen, 1826, by monotypy. ern Desert: Wadi Rishrash, Wadi Wirak. Lower Nile Valley Turanogonia Rohdendorf, 1924: 228. Type species: and Delta: Giza, Helwan, Kerdassa, Tanta. Sinai: El-Arish, Ein Turanogonia smimovi Rohdendorf, 1924 (= Gonia Gedeirat,Ein Moweileh,Gebel Moussa,WadiEl-Lega,Wadi chinensis Wiedemann, 1824), by monotypy. Firan. Western Desert: Siwa Oasis. Upper Nile Valley: Asswan. Asiogonia Rohdendorf, 1928: 98. Type species: Dates of collection: February to October. Asiogonia asiatica Rohdendorf, 1928, by monotypy. Gonia ornata Meigen, 1826 Chrysocerogonia Rohdendorf, 1928: 98 (as subgenus of Gonia ornata Meigen, 1826: 3. Syntypes, males Salmacia Meigen, 1800). Type species: Salmacia (MNHN). France (Lyon). (Chrysocerogonia) ussuriensis Rohdendorf, 1928,by Gonia vittata Meigen, 1826: 3. Germany. monotypy. Gonia nervosa Meigen, 1826: 4. Germany (Hamburg). Eremogonia Rohdendorf, 1928: 98 (as subgenus of Rhedia vernalis Robineau-Desvoidy, 1830: 74. France. Salmacia Meigen, 1800). Type species: Salmacia Rhedia diversa Robineau-Desvoidy, 1830: 77. France. (Eremogonia) desertorum Rohdendorf, 1928,by monotypy. Gonia lateralis Zeller, 1842: 843. Poland (Glogow). Gonia atra Meigen, 1826 Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Gonia atra Meigen, 1826: 7. Unspecified type(s) Turkmenistan, Uzbekistan), China, Egypt, Europe (all), M. (MNHN). France. East, Mongolia, Russia (W. Russia, W. Siberia, S. Far East), Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Transcaucasia. Turkmenistan, Uzbekistan), China, Egypt, Europe, Egyptian localities: Common throughout the country, Kazakhstan, Mongolia, Russia (Siberia), Transcaucasia. particularly in: Lower Nile Valley and Delta: Abu Egyptian localities: Fayoum:? . Lower Nile Valley and Delta: Rawash, Tanta. Upper Nile Valley: Qena. El-Marg, Ezbet El-Nakhl, Kafr Hakim, Kerdassa, Sentrees. Dates of collection: February to August. Dates of collection: February to October. Genus MENDELSSOHNIA Kugler Gonia bimaculata Wiedemann, 1819 Mendelssohnia Kugler 1971: 69. Type species: Gonia bimaculata Wiedemann, 1819: 25. Syntypes, Mendelssohnia sinaica Kugler, 1971, by original females (NHMW, ZMUC). South Africa (Western Cape: designation. Cape of Good Hope). Mendelssohnia sinaica Kugler, 1971 Gonia cilipeda Rondani, 1859: 35. Italy (Parma and Mendelssohnia sinaica Kugler, 1971: 70. Holotype Malta). male (TAU). Egypt (Sinai: Wadi El-Sheikh). Gonia incerta Bigot, 1888: 10. (Tunisia. Tunis). Distribution: Egypt, Israel. Salmacia cabrerae Mesnil, 1956: 523. Canary Is. Egyptian localities: Sinai: Wadi El Sheikh. (Hierro). Dates of collection: Unknown. Distribution: AF: Widespread (excl. W. AF). OR: Orien. Genus NEALSOMYIA Mesnil China[seeO’Hara and Cerretti 2016: 19]. PA: Canary Is., C. Nealsomyia Mesnil, 1939: 31. Type species: Exorista Asia (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), (Alsomyia) triseriella Villeneuve, 1929, by original Europe, Israel, Madeira, North Africa, Pal. China [see designation. O’Hara and Cerretti 2016: 19], Transcaucasia. Nealsomyia triseriella (Villeneuve, 1929) Egyptian localities: CoastalStrip: Abu Qir. Eastern Exorista triseriella Villeneuve, 1929c: 185. Holotype Desert: Gebel Asfar, Ismailia, Wadi Hoff. Fayoum: male (CNC). Egypt (Wadi Hoff: nr. Helwan). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 9 of 19 Distribution: PA: Egypt, Israel. Ramonella Kugler, 1980b:67(nomen novum for Egyptian localities: Eastern Desert: Wadi Hoff. Ramona Kugler, 1980). Dates of collection: April to August. Ramonella mesnili (Kugler, 1980) Genus PSEUDOGONIA Brauer and Bergenstamm Ramona mesnili Kugler, 1980a: 41. Holotype male Pseudogonia Brauer and Bergenstamm, 1889: 100. (TAU). Israel (Negev: Ramon). Type species: Gonia cinerascens Rondani, 1859 Distribution: AF: Yemen. PA: Canary Is., Egypt, (= Tachina rufifrons Wiedemann, 1830), by monotypy. Turkey, Israel. Gaediogonia Townsend, 1927: 71. Type species: Egyptian localities: Lower Nile Valley and Delta: El- Gaediogonia jacobsoni Townsend, 1927 (= Tachina Marg. Sinai: Wadi Taibe, Wadi Watir. rufifrons Wiedemann, 1830), by original designation. Dates of collection: May to September. Pseudogonia fasciata (Wiedemann, 1819) Gonia fasciata Wiedemann, 1819: 25. Syntypes, Tribe WINTHEMIINI females (ZMUC). South Africa (Western Cape: Cape of Genus NEMORILLA Rondani Good Hope). Nemorilla Rondani, 1856: 66. Type species: Tachina Rhedia capensis Robineau-Desvoidy, 1830: 77. South maculosa Meigen, 1824, by original designation. Africa (Western Cape: Cape of Good Hope). Nemorilla floralis (Fallén, 1810) Reaumuria lalandii Robineau-Desvoidy, 1830: 80. Tachina floralis Fallén, 1810: 287. Holotype male South Africa (Western Cape: Cape of Good Hope). (NHRS). Sweden. Distribution: AF: South Africa, Zimbabwe. PA: Tachina notabilis Meigen, 1824: 266. Sweden. Andorra, Canary Is., Egypt, Portugal (including Azores, Tachina angustipennis Meigen, 1824: 333. W. Europe. excluding Madeira), Spain. Tachina pabulina Meigen, 1824: 358. Germany Egyptian localities and dates of collection are not (Hamburg). precisely known. Recorded from Egypt by Steyskal and Exorista laticella Macquart, 1850: 373. France El-Bialy 1967: 49). (Lestrem). Pseudogonia rufifrons (Wiedemann, 1830) Exorista arrogans Macquart, 1850: 374. France Tachina rufifrons Wiedemann, 1830: 318. Lectotype (Lestrem). female (ZMUC). China. Tachina intersita Walker, 1853: 72. England. Latreillia lalandii Robineau-Desvoidy, 1830: l06. South Nemorilla amica Rondani, 1859: 102. Italy (Parma). Africa (Western Cape: Cape of Good Hope). Aubaea campestris Robineau-Desvoidy, 1863: 186. Gonia cinerascens Rondani, 1859: 34. Italy (Parma). France. Gonia munroi Curran, 1927b: 339. Tanzania. Pitthaea nebulosa Robineau-Desvoidy, 1863: 189. Gonia ritchiei Cuthbertson and Munro, 1941:109. France. Nomen nudum. Distribution: PA: Egypt, Europe, Russia, Japan, North Gonia munroi curran, 1927b: 339) Tanzania (Tanganyika) Korea. and South Africa. Egyptian localities: Lower Nile Valley and Delta: Distribution: AF: Widespread, including Cape Verde, Bahtim, Ezbet El-Nakhl. Sinai: El-Arish. Nigeria, South Africa, Tanzania, U.A. Emirates, Yemen. Dates of collection: April to June. AU: Australia, Hawaii, Melanesia, N. Australasian. OR: India, Indonesia, Malaysia, Myanmar, Orien. China [see Subfamily PHASIINAE O’Hara and Cerretti 2016: 19], Pakistan, Philippines, Tribe CYLINDROMYIINI Ryukyu Is., Taiwan, Thailand. PA: C. Asia (Kyrgyzstan, Genus BESSERIA Robineau-Desvoidy Tajikistan, Turkmenistan, Uzbekistan), Egypt, all Europe Besseria Robineau-Desvoidy, 1830: 232. Type species: (except British Is.), Japan, Kazakhstan, Korea, M. East Besseria reflexa Robineau-Desvoidy, 1830, by monotypy. (Israel), Mongolia, Pal. China [see O’Hara and Cerretti Apostrophus Loew, 1871: 310, 311. Type species: 2016: 19], Russia, Transcaucasia. Apostrophus suspectus Loew, 1871 (=Actia zonaria Egyptian localities: Coastal Strip: Mariout, Nuzha. Loew, 1847), by subsequent designation of Coquillett Eastern Desert: Wadi Hoff. Gebel Elba: Wadi Edeib. (1910: 509). Lower Nile Valley and Delta: Abu Rawash. Sinai: Wadi Apostrophusia Townsend, 1933: 454. Type species: El-Tih. Apostrophus anthophilus Loew, 1871, by original Dates of collection: January to July. designation. Genus RAMONELLA Kugler Besseria anthophila (Loew, 1871) Ramona Kugler, 1980a: 40 (junior homonym of Apostrophus anthophilus Loew, 1871: 310. Type(s), Ramona Casey, 1886). Type species: Ramona mesnili male(s), not located. Germany (Bavaria: Munich- Kugler, 1980, by original designation. Geiselgasteig). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 10 of 19 Melia forcipata Bigot, 1881: 368. France (Hautes- Dionomelia Kugler, 1978b: 346. Type species: Alpes: Queyras, Abries). Dionomelia hennigi Kugler, 1978, by original designation. Distribution: NE: Alberta, Northwest Territories. PA: Dionomelia hennigi Kugler, 1978 Egypt, Europe, Mongolia. Dionomelia hennigi Kugler, 1978b: 346. Holotype Egyptian localities: Coastal Strip: Dekhila, Mariout. female (TAU). Israel (Dead Sea area: Ein Boqeq). Dates of collection: June to August. Distribution: AF: U.A. Emirates. PA: Andorra, Egypt, Genus CYLINDROMYIA Meigen Israel, Portugal (including Azores, excluding Madeira), Cylindromyia Meigen, 1803: 279. Type species: Spain. Musca brassicaria Fabricius, 1775, by monotypy. Egyptian localities: Sinai. Ocyptera Latreille, 1804: 195. Type species: Musca Dates of collection: April to September. brassicaria Fabricius, 1775, by subsequent designation of Genus LEUCOSTOMA Meigen Curtis (1837: 629). Leucostoma Meigen, 1803: 279. Type species: Exogaster Rondani, 1856: 78. Type species: Exogaster Ocyptera simplex Fallén, 1815, by subsequent monotypy carinatus Rondani, 1856 (= Ocyptera rufifrons Loew, of Meigen (1824: 234). 1844), by original designation. Leucostoma engeddense Kugler, 1966 Ocypterula Rondani, 1856: 78. Type species: Ocyptera Leucostoma engeddense Kugler, 1966: 177. Holotype pusilla Meigen, 1824, by original designation. female (TAU). Israel (Dead Sea area: Ein Gedi). Plesiocyptera Brauer and Bergenstamm, 1893: 56. Type Distribution: AF: South Africa, U.A. Emirates. PA: species: Ocyptera bicolor Wiedemann, 1819,by Algeria, Canary Is., Egypt, Israel, Madeira, NW Africa, monotypy. Spain, Turkey. Conopisoma Speiser, 1910: 144. Type species: Egyptian localities: Lower Nile Valley and Delta: Minia Conopisoma miraculum Speiser, 1910, by original El-Qamh, Tanta. designation. Dates of collection: February to May. Formicocyptera Townsend, 1933: 451. Type species: Leucostoma obsidianum (Wiedemann, 1830) Ocyptera atrata Fabricius, 1805, by original designation. Tachina obsidiana Wiedemann, 1830: 341. Lectotype Cylindromyia intermedia (Meigen, 1824) female (SMF). Sudan (Nuba [as “Nubien”]). Ocyptera intermedia Meigen, 1824: 212. Holotype Leucostoma marismortui Kugler, 1966: 179. Israel male (MNHN). ?Europe. (Dead Sea area: Ein Gedi). Ocyptera scalaris Loew, 1844: 240. Austria (Vienna). Distribution: AF: Sudan, Yemen. PA: Canary Is., Egypt, Ocyptera excisa Loew, 1845: 171. Romania (Mehadia). Europe, Israel, Malta, Saudi Arabia. Ocyptera excisa var. rufiventris Strobl, 1906: 338. Spain Egyptian localities: Upper Egypt: Asswan. (Villa Rutis). Dates of collection: Unknown. Ocyptera reinigi Enderlein, 1934: 132. Tajikistan (Pamir: Karasu). Tribe PHASIINI Distribution: Nearctic: Widespread. PA: C. Asia Genus PHASIA Latreille (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), Phasia Latreille, 1804: 195. Type species: Conops China, Egypt, Europe, Iran, Israel, Mongolia, Russia, subcoleoptratus Linnaeus, 1767, by subsequent Tadzhikistan, Transcaucasia, Ukraine, Uzbekistan. monotypy of Latreille (1805: 379). Egyptian localities: Coastal Strip: Alexandria, Mariout, Alophora Robineau-Desvoidy, 1830: 293. Type species: Max, Ramleh. Upper Nile Valley: Beni Suef. Syrphus hemipterus Fabricius, 1794, by subsequent des- Dates of collection: April to September. ignation of Robineau-Desvoidy (1863b: 226, as “Thereva Cylindromyia rufipes (Meigen, 1824) hemiptera de Fabricius”). Ocyptera rufipes Meigen, 1824: 215. Lectotype male Hyalomya Robineau-Desvoidy, 1830: 298. Type spe- (MNHN). France. cies: Phasia semicinerea Meigen, 1824 (=Phasia pusilla Distribution: AF: U.A. Emirates. OR: India, Pakistan. Meigen, 1824), by subsequent designation of Westwood PA: Egypt, all Europe (except British Is., Scand.), (1840: 140). Pakistan, Russia (W. Russia), Saudi Arabia, Phorantha Rondani, 1861b: 209. Type species: Transcaucasia, Ukraine. Phorantha musciformis Rondani, 1861 (= Conops Egyptian localities: Coastal Strip: Alexandria. Lower subcoleoptratus Linnaeus, 1767), by monotypy. Nile Valley and Delta: Gebel Asfar, Kubba. Paralophora Girschner, 1887: 412 (as subgenus of Dates of collection: October to February. Alophora Robineau-Desvoidy, 1830). Type species: Phasia pusilla Meigen, 1824, by monotypy. Tribe LEUCOSTOMATINI Alophorella Townsend, 1912: 45. Type species: Genus DIONOMELIA Kugler Thereva obesa Fabricius, 1798, by original designation. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 11 of 19 Phasia pusilla Meigen, 1824 Tachinomima Brauer and Bergenstamm, 1891: 383. Phasia pusilla Meigen, 1824: 198. Lectotype male Type species: Tachinomima expetens Brauer and (CNC). Germany (Stolberg). Bergenstamm, 1891 (= Micropalpus longirostris Phasia semicinerea Meigen, 1824: 199. Germany Macquart, 1844), by monotypy. (Stolberg). Hecatoepalpus Townsend, 1933: 467. Type species: Hyalomya carbonaria Robineau-Desvoidy, 1830: 300. Micropalpus prohecate Speiser, 1910, by original France (St. Sauveur). designation. Hyalomya corinna Robineau-Desvoidy, 1830: 301. Micropalpinus Enderlein, 1937: 441. Type species: France. Micropalpus pallidus Jaennicke, 1867, by original Phasia vitripennis Zetterstedt, 1859: 6169. Sweden designation. (Skane). Gymmantia Enderlein, 1937: 441. Type species: Hyalomya chorea Robineau-Desvoidy, 1863: 266. Micropalpus alboscutellatus Speiser, 1910, by original France. designation. Distribution: PA: C. Asia (Kyrgyzstan, Tajikistan, Linnaemya comta (Fallén, 1810) Turkmenistan, Uzbekistan), China, Egypt, Europe (all), Tachina comta Fallén, 1810: 277. Unspecified type(s), Israel, Japan, Kazakhstan, Mongolia, Russia, Transcaucasia. females (NHRS) Sweden. Egyptian localities and dates of collection are not Tachina compta Meigen, 1824: 262. Unjustified precisely known. emendation of comta Fallén. Tachina fulgens Meigen, 1824: 259. Austria, France Subfamily TACHININAE and Germany. Tribe ERNESTIINI Linnaemya heraclei Robineau-Desvoidy, 1830: 53. Genus EURITHIA Robineau-Desvoidy France (Paris). Erigone Robineau-Desvoidy, 1830: 65 (junior homo- Linnaemya analis Robineau-Desvoidy, 1830: 54. nym of Erigone Audouin, 1826). Type species: Erigone France (Angers). anthophila Robineau-Desvoidy, 1830, by subsequent Linnaemya aestivalis Robineau-Desvoidy, 1830: 54. designation of Townsend (1932: 42). France. Eurithia Robineau-Desvoidy, 1844: 24. Type species: Linnaemya borealis Robineau-Desvoidy, 1830: 54. Erigone puparum Robineau-Desvoidy, 1830 (= Tachina France. caesia Fallén, 1810), by monotypy. Bonnetia longipes Robineau-Desvoidy, 1830: 55. Eurithia castellana (Strobl, 1906) France. Erigone castellana Strobl, 1906: 338. Holotype male Bonnetia oenanthis Robineau-Desvoidy, 1830: 56. (NMBA). Spain (Madrid). Holotype may be lost, see France (Paris). O’Hara et al. (2009). Micropalpus humeralis Robineau-Desvoidy, 1863: 119. Distribution: PA: Albania, Bosnia and Herzegovina, New name for Bonnetia longipes Robineau-Desvoidy, 1830. Bulgaria, China, Croatia, Egypt, Greece, Israel, Distribution: AF: Sudan. NE: Widespread. OR: India, Montenegro, Macedonia, Serbia, Slovenia, Transcaucasia. Nepal. PA: C. Asia (Kyrgyzstan, Tajikistan, Egyptian localities: Coastal Strip: El-Burg, Mariout. Turkmenistan, Uzbekistan), China, Egypt, Europe (all), Eastern Desert: Ismailia. Fayoum: ?. Lower Nile Valley Kazakhstan, Israel, Mongolia, Russia (W. Russia, W. and Delta: Abu Rawash. Upper Nile Valley: Asswan. Siberia, E. Siberia, S. Far East), Taiwan, Transcaucasia. Dates of collection: January to October. Egyptian localities: Eastern Desert: Wadi Garawi, Wadi Genus LINNAEMYA Robineau-Desvoidy Rishrash. Gebel Elba: ?. Lower Nile Valley and Delta: Linnaemya Robineau-Desvoidy, 1830: 52. Type spe- Mansouriya. cies: Linnaemya silvestris Robineau-Desvoidy, 1830 Dates of collection: January to March. (= Tachina vulpina Fallén, 1810), by subsequent designa- Linnaemya latigena Kugler, 1977 tion of Robineau-Desvoidy (1863a: 131). Linnaemyia latigena Kugler, 1977: 3. Holotype male Bonellia Robineau-Desvoidy, 1830: 56 (junior homo- (TAU). Egypt (Sinai: Bir Gifgafa [as “Refidim”]). nym of Bonellia Rolando, 1822). Type species: Bonellia Distribution: AF: U.A. Emirates. PA: Egypt, Israel. tessellans Robineau-Desvoidy, 1830, by subsequent des- Egyptian localities: Sinai: Bir Gifgafa, Mitla, Wadi ignation of Townsend (1916: 6). Firan. Elachipalpus Rondani, 1850: 169. Type species: Dates of collection: April. Micropalpus longirostris Macquart, 1845 (junior primary Linnaemya petiolata Kugler, 1971 homonym of Micropalpus longirostris Macquart, 1844; Linnaemya petiolata Kugler, 1971: 80. Unspecified = Elachipalpus rondanii Townsend, 1916), by original type(s), male(s) (TAU). Egypt (South Sinai: Abu designation. Marawath). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 12 of 19 Distribution: PA: Egypt. Egyptian localities. Coastal Strip: Alexandria, Amrya. Egyptian localities: Sinai: Abu Marawath. Lower Nile Valley and Delta: Abu Rawash, Ain Shams, Dates of collection: April to August. Beni Meggon, Cairo, Dokki, El-Baragil, Ezbet El-Nakhl, Linnaemya soror Zimin, 1954 Giza, Kafr Hakim, Kerdassa, Mansouriya, Maadi, Nahya, Linnaemyia soror Zimin, 1954: 266. Holotype male Shubra, Sinai: El-Arish. Sinai: El-Arish. Upper Nile Val- (ZIN). Tajikistan (Gorno-Badakhshan: Khorugh). ley: Assiout, Asswan. Distribution: PA: Canary Is., C. Asia (Kyrgyzstan, Dates of collection. Throughout the year. Tajikistan, Turkmenistan, Uzbekistan), China, Egypt, W. Genus PLESINA Meigen Europe (Andorra, Portugal (including Azores, excluding Plesina Meigen, 1838: 214. Type species: Tachina Madeira), Spain), S. Europe (Albania, Bosnia and phalerata Meigen, 1824, by monotypy. Herzegovina, Bulgaria, Croatia, Greece, Montenegro, Xanthopetia Townsend, 1933: 452. Type species: Macedonia, Serbia, Slovenia), Israel, North Africa, Tachina fascipennis Wiedemann, 1830, by original Russia, Transcaucasia, Turkmenia, Uzbekistan. OR: designation. India, Nepal. Kugleria Verbeke, 1970: 299 (junior homonym of Egyptian localities: Pyramids. Kugleria Bouwman, 1938). Type species: Plesina Dates of collection: September. fascipennis claripennis Mesnil, by monotypy. Plesina claripennis Mesnil, 1953 Plesina claripennis Mesnil, 1953b: 166. Holotype (not Tribe GRAPHOGASTRINI located). Israel (Jerusalem). Genus PHYTOMYPTERA Rondani Distribution: Egypt, Israel. Phytomyptera Rondani, 1845: 32, 33. Type species: Egyptian localities: Sinai. Phytomyptera nitidiventris Rondani, 1845 (= Tachina Dates of collection: ? nigrina Meigen, 1824), by monotypy. Plesina deserticola Kugler, 1978 Elfia Robineau-Desvoidy, 1850: 190. Type species: Plesina deserticola Kugler, 1978a: 93. Holotype male Actia cingulata Robineau-Desvoidy, 1830, by subsequent (TAU). Egypt (Sinai: Wadi Shag). designation of Robineau-Desvoidy (1863a: 672). Distribution: PA: Egypt, Israel. Phytomyptera lacteipennis Villeneuve, 1934 Egyptian localities: Sinai: Wadi Shag (South Sinai). Phytomyptera lacteipennis Villeneuve, 1934b: 71. Dates of collection: ? Lectotype female (CNC). Egypt (Suez). Genus ROSSIMYIOPS Mesnil Distribution: AF: U.A. Emirates. PA: Egypt, Europe, Rossimyiops Mesnil, 1953a: 145. Type species: Israel, Libya, Mongolia, Russia (W. Russia). Rossimyiops whiteheadi Mesnil, 1953, by monotypy. Egyptian localities: Eastern Desert: Suez Road. Mesnilomyia Kugler, 1972: 103. Type species: Dates of collection: March. Mesnilomyia magnifica Kugler, 1972, by original designation. Tribe MINTHOINI Persedea Richter, 2001: 25. Type species: Persedea Genus MINTHO Robineau-Desvoidy exquisita Richter, 2001, by original designation. Mintho Robineau-Desvoidy, 1830: 216. Type species: Rossimyiops achilleae (Kugler, 1972) Musca compressa Fabricius, 1787, by subsequent desig- Mesnilomyia achilleae Kugler, 1972: 107. Holotype nation of Rondani (1856: 79). male (TAU). Israel (‘Arad). Mintho compressa (Fabricius, 1787) Distribution: PA: Egypt, Israel. Musca compressa Fabricius, 1787: 346. Unspecified Egyptian localities: Sinai: St. Catherine, Wadi El- type(s) (ZMUC). Spain. Arbaein, Wadi Shag. Mintho capensis Robineau-Desvoidy, 1830: 217. South Dates of collection: July. Africa (Western Cape: Cape of Good Hope). Rossimyiops magnificus (Kugler, 1972) Tachina isis Wiedemann, 1830: 304. Egypt. Mesnilomyia magnifica Kugler, 1972: 105. Holotype Ocyptera vittata Costa, 1844: 94. Italy (Cervinara and male (TAU). Israel (‘Arad). Guagnano). Distribution: PA: Egypt, Iraq, Israel. Sericocera algira Macquart, 1849: 480. Algeria. Egyptian localities: Sinai: Tarfat El-Kadarin, Wadi Dexia thala Walker, 1849: 845. Morocco (Tangier). Nazeb, Wadi Sa’al, Nueiba’. Dexia isapis Walker, 1849: 848. Egypt. Dates of collection: April to September. Distribution: AF: Widespread, including D.R. Congo, Eritrea, Kenya, Nigeria, Somalia, South Africa, Sudan, Tribe SIPHONINI Tanzania, Yemen. PA: Algeria, Canary Is., Egypt, Europe, Genus ACTIA Robineau-Desvoidy Israel, Libya, Morocco, Transcaucasia, Tunisia. El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 13 of 19 Actia Robineau-Desvoidy, 1830: 85. Type species: Roe- Taiwan, Thailand. PA: Japan, Israel, North Africa, P. selia lamia Meigen, 1838, by designation under the plen- China. AU: Australia, Melanesia, Micronesia, N. ary powers of ICZN. Australasian. Gymnophthalma Lioy, 1864: 1341. Type species: Egyptian localities: Lower Nile Valley and Delta: Giza, Tachina crassicornis Meigen, 1824, by monotypy. Sidi Ghazi, Torah. Upper Nile Valley: Beni Morr (Assiout). Gymnopareia Brauer and Bergenstamm, 1889:103. Dates of collection: May to August. Type species: Tachina crassicornis Meigen, 1824,by Peribaea palaestina (Villeneuve, 1934) monotypy. Actia palaestina Villeneuve, 1934a: 57. Holotype Actia crassicornis (Meigen, 1824) female (SMNS). Israel, (Rehovot [as “Rehoboth”]). Tachina crassicornis Meigen, 1824: 351. Syntypes, Actia alipes Villeneuve, 1942: 134. Egypt (Asswan [as males and females (MNHN). Probably Germany (Stolberg). “Assuan”]). See O’Hara (2009). Distribution: AF: U.A. E., Yemen. PA: C. Asia Thryptocera bicolor Macquart, 1834: 312. France (Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), (Lille). China, Egypt, Iran, Israel. Thryptocera flavipalpis Macquart, 1848: 135. Egyptian localities and dates of collection are not Switzerland (Ütliberg). precisely known. Thryptocera nigripalpis Robineau-Desvoidy, 1851: 182. Genus SIPHONA Meigen France. Crocuta Meigen, 1800: 39. Name suppressed by ICZN. Thryptocera palpalis Rondani, 1859: 14. Italy (Parma). Siphona Meigen, 1803: 281. Type species: Musca Thryptocera claripennis Robineau-Desvoidy, 1863: 716 geniculata De Geer, 1776, by designation under the France, plenary powers of ICZN. Thryptocera scutellaris Rondani, 1865: 195. Italy Subgenus SIPHONA Meigen, 1803 —Nominotypical (Parma). subgenus. Distribution: PA: China, Egypt, Europe (all), Japan, Siphona (Siphona) efflatouni Mesnil, 1960 Kazakhstan, Mongolia, Russia, Transcaucasia. Siphona efflatouni Mesnil, 1960: 188. Holotype female Egyptian localities: Coastal Strip: Alexandria. Eastern (CNC). Egypt (Mariout). Desert: Ismailia. Lower Nile Valley and Delta: Abu Distribution: PA: Egypt, Israel. Rawash, Dokki, Giza, Kerdassa. Sinai: El-Arish. Upper Egyptian localities: Coastal Strip: Mariout. Sinai: El-Arish. Nile Valley: Assiout. Dates of collection: March to August. Dates of collection: April to September. Genus PERIBAEA Robineau-Desvoidy Tribe TACHININI Herbstia Robineau-Desvoidy, 1851: 184 (junior homo- Genus PELETERIA Robineau-Desvoidy nym of Herbstia Edwards, 1834). Type species: Herbstia Peleteria Robineau-Desvoidy, 1830: 39. Type species: tibialis Robineau-Desvoidy, 1851, by monotypy. Peleteria abdominalis Robineau-Desvoidy, 1830, by sub- Peribaea Robineau-Desvoidy, 1863a: 720. Type spe- sequent designation of Coquillett (1910: 586). cies: Peribaea apicalis Robineau-Desvoidy, 1863 (= Herb- Cuphocera Macquart, 1845: 267. Type species: stia tibialis Robineau-Desvoidy, 1851), by subsequent Micropalpus ruficornis Macquart, 1835, by original designation of Coquillett (1910: 587). designation. Strobliomyia Townsend, 1926a: 31. Type species: Acuphocera Townsend, 1926b: 37. Type species: Tryptocera fissicornis Strobl, 1910 (as “Thryptocera Acuphocera sumatrensis Townsend, 1926 (= Tachina fissicornis”)(= Thryptocera setinervis Thomson, 1869), iavana Wiedemann, 1819), by original designation. by original designation. Pleropeleteria Villeneuve, 1916: 470. Type species: Peribaea orbata (Wiedemann, 1830) Dejeania (Pleropeleteria) peringueyi Villeneuve, 1916 Tachina orbata Wiedemann, 1830: 336. Neotype (= Tachina lithanthrax Wiedemann, 1830), by monotypy. female (BMNH). India (Assam: Azra). Peletieriana Mesnil, 1970a: 951 (as subgenus of Gymnopareia (Actia) aegyptia Villeneuve, 1913a: 508. Peleteria Robineau-Desvoidy, 1830,as “Peletieria”). Type Egypt (Qaliûb). species: Echinomyia rustica Karsch, 1886, by original Actia nigripes Curran, 1927a: 6. D.R. Congo (Bas- designation. Congo: Boma). Peleteria meridionalis (Robineau-Desvoidy, 1830) Strobliomyia sororcula Mesnil, 1954: 16. D.R. Congo Faurella meridionalis Robineau-Desvoidy, 1830: 41. (Nord-Kivu: Rutshuru). Type(s) not located. France. Distribution: AF: Widespread. OR: India, Indonesia, Echinomyia ruficeps Macquart, 1845: 263. Italy. Orien. China [see O’Hara and Cerretti 2016: 19], Peleteria vernalis Robineau-Desvoidy, 1863: 617. Malaysia, Myanmar, Philippines, Ryukyu Is., Sri Lanka, France (Hyeres). El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 14 of 19 Peleteria pedemontana Robineau-Desvoidy, 1863: 619. Genus UCLESIA Girschner Italy. Peleteria algira Robineau-Desvoidy, 1863: 621. Algeria. – Uclesia petiolata (Villeneuve, 1929) Peletieria capitata Zimin, 1961: 265. Turkmenistan (Chandyr Valley). Genus WAGNERIA Robineau-Desvoidy Distribution: PA: Algeria, Egypt, Isael, Italy, Kirghizia, Russia, Tadzhikistan, Transcaucasia, Uzbekistan. – Wagneria cunctans (Meigen, 1824) Egyptian localities: Coastal Strip: Alexandria, Mariout. Fayoum: ?. Lower Nile Valley and Delta: Kerdassa, Dokki, Subfamily EXORISTINAE Maadi. Sinai: El-Arish. Upper Nile Valley: Assiout. Tribe ERYCIINI Dates of collection: November to April. Genus AMELIBAEA Mesnil List of doubtful taxa (Nomina Dubia) described from Egypt: (see Herting and Dely-Draskovits 1993) – Amelibaea ?signifera (Villeneuve, 1929) – Catabucentes abdominalis Villeneuve, 1934b: 72. Genus APLOMYA Robineau-Desvoidy Egypt (Lower Egypt). – Eurigaster incisus Walker, 1871:340.Egypt (HotTamanib). – Aplomya confinis (Fallén, 1820) – Eurigaster leucomelas Walker, 1871: 339. Egypt. – Aplomya metallica (Wiedemann, 1824) – Frontina latifascia Walker, 1871: 340. Egypt (Wadi Genneh). Genus CADURCIELLA Villeneuve – Reaumuria olivieri Robineau-Desvoidy, 1830:82.Egypt. – Reaumuria virescens Robineau-Desvoidy, 1863b: 741. – Cadurciella tritaeniata (Rondani, 1859) Egypt. Genus CESTONIA Rondani Conclusion As a final conclusion, this catalogue treats a total of 72 – Cestonia cineraria Rondani, 1861 tachinid species belonging to 42 genera, 15 tribes and 4 – Cestonia rutilans Villeneuve, 1929 subfamilies in Egypt. Synonymies, type localities, type depositories, world distributions by biogeographic Genus CESTONIONERVA Villeneuve realm(s) and country, Egyptian localities and dates of collection are provided for all Egyptian taxa. – Cestonionerva petiolata (Villeneuve, 1910) Appendix Genus DRINO Robineau-Desvoidy Checklist of the Tachinidae of Egypt Subgenus PALEXORISTA Townsend Subfamily DEXIINAE Tribe DEXIINI – Drino (Palexorista) gilva (Hartig, 1838) Genus TRIXICEPS Villeneuve – Drino (Palexorista) imberbis (Wiedemann, 1830) – Drino (Palexorista) inconspicua (Meigen, 1830) – Trixiceps magnipalpis (Bezzi, 1922) Subgenus ZYGOBOTHRIA Mik Tribe VORIINI Genus HYPOVORIA Villeneuve – Drino (Zygobothria) atropivora (Robineau-Desvoidy, 1830) – Hypovoria hilaris (Villeneuve, 1913) Genus THECOCARCELIA Townsend Genus NANOPLAGIA Villeneuve – Thecocarcelia acutangulata (Macquart, 1851) – Nanoplagia sinaica (Villeneuve, 1909) Tribe ETHILLINI Genus ATYLOMYIA Brauer Genus PERISCEPSIA Gistel Subgenus PERISCEPSIA Gistel – Atylomyia albifrons Villeneuve, 1911 – Periscepsia (Periscepsia) handlirschi (Brauer and von Tribe EXORISTINI Bergenstamm, 1891) Genus ALLOPROSOPAEA Villeneuve El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 15 of 19 – Alloprosopaea efflatouni Villeneuve, 1923 Genus NEALSOMYIA Mesnil Genus CHAETORIA Becker – Nealsomyia triseriella (Villeneuve, 1929) – Chaetoria stylata Becker, 1908 Genus PSEUDOGONIA Brauer and Bergenstamm Genus CHETOGENA Rondani – Pseudogonia fasciata (Wiedemann, 1819) – Pseudogonia rufifrons (Wiedemann, 1830) – Chetogena aegyptiaca (Villeneuve, 1923) – Chetogena cercosa (Kugler, 1980) Genus RAMONELLA Kugler – Chetogena sinaica (Villeneuve, 1909) – Ramonella mesnili (Kugler, 1980) Genus EXORISTA Meigen. Subgenus ADENIA Robineau-Desvoidy Tribe WINTHEMIINI Genus NEMORILLA Rondani – Exorista (Adenia) rustica (Fallén, 1810) – Nemorilla floralis (Fallén, 1810) Subgenus EXORISTA Meigen Subfamily PHASIINAE – Exorista (Exorista) fasciata (Fallén, 1820) Tribe CYLINDROMYIINI – Exorista (Exorista) larvarum (Linnaeus, 1758) Genus BESSERIA Robineau-Desvoidy Subgenus PODOTACHINA Brauer and Bergenstamm – Besseria anthophila (Loew, 1871) – Exorista (Podotachina) sorbillans (Wiedemann, 1830) Genus CYLINDROMYIA Meigen Subgenus PTILOTACHINA Brauer and Bergenstamm – Cylindromyia intermedia (Meigen, 1824) – Cylindromyia rufipes (Meigen, 1824) – Exorista (Ptilotachina) rutilans Mesnil, 1970 – Exorista (Ptilotachina) xanthaspis (Wiedemann, 1830) Tribe LEUCOSTOMATINI Genus DIONOMELIA Kugler Genus MACULOSALIA Mesnil – Dionomelia hennigi Kugler, 1978 – Maculosalia maculosa (Villeneuve, 1909) Genus LEUCOSTOMA Meigen Tribe GONIINI Genus BLEPHARELLA Macquart – Leucostoma engeddense Kugler, 1966 – Leucostoma obsidianum (Wiedemann, 1830) – Blepharella versatilis (Villeneuve, 1910) Tribe PHASIINI Genus CLEMELIS Robineau-Desvoidy Genus PHASIA Latreille – Clemelis apicalis Villeneuve, 1923 – Phasia pusilla Meigen, 1824 Genus GONIA Meigen Subfamily TACHININAE Tribe ERNESTIINI – Gonia atra Meigen, 1826 Genus EURITHIA Robineau-Desvoidy – Gonia bimaculata Wiedemann, 1819 – Gonia capitata (De Geer, 1776) – Eurithia castellana (Strobl, 1906) – Gonia ornata Meigen, 1826 Genus LINNAEMYA Robineau-Desvoidy Genus MENDELSSOHNIA Kugler – Linnaemya comta (Fallén, 1810) – Mendelssohnia sinaica Kugler, 1971 – Linnaemya latigena Kugler, 1977 El-Hawagry Egyptian Journal of Biological Pest Control (2018) 28:46 Page 16 of 19 – Linnaemya petiolata Kugler, 1971 London, London, United Kingdom; MHNL: Musée d’Histoire Naturelle de Lille, Lille, France; MNHN: Muséum National d’Histoire Naturelle, Paris, France; – Linnaemya soror Zimin, 1954 MRSN: Museo Regionale di Scienze Naturali, Torino [Turin], Italy; MZF: Museo Zoologico “La Specola”, Firenze [Florence], Italy; MZLU: Museum of Zoology, Tribe GRAPHOGASTRINI Lund University, Lund, Sweden; NHMW: Naturhistorisches Museum Wien, Wien [Vienna], Austria; NHRS: Naturhistoriska riksmuseet [Swedish Museum of Genus PHYTOMYPTERA Rondani Natural History], Stockholm, Sweden; NMBA: Naturhistorisches Museum der Benediktiner-Abtei Admont, Admont, Austria; PPDD: Ministry of Agriculture – Phytomyptera lacteipennis Villeneuve, 1934 Collection, Plant Protection Research Institute, Dokki, Giza, Egypt; SMF: Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany; SMNS: Staatliches Museum für Naturkunde, Stuttgart, Germany; Tribe MINTHOINI TAU: Tel Aviv University, Tel Aviv, Israel; USNM: National Museum of Natural History [formerly United States National Museum], Smithsonian Institution, Genus MINTHO Robineau-Desvoidy Washington, District of Columbia, USA; ZIN: Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia; ZMHB: Museum für Naturkunde – Mintho compressa (Fabricius, 1787) der Humboldt-Universität zu Berlin, Berlin, Germany; ZMUC: Zoological Museum, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark; ZMUH: Universität von Hamburg, Zoologisches Genus PLESINA Meigen Institut und Zoologisches Museum, Hamburg, Germany Biogeographic Realm abbreviations used throughout the catalogue – Plesina claripennis Mesnil, 1953 AF: Afrotropical; AU: Australasian; NE: Nearctic; NT: Neotropical; OR: Oriental; – Plesina deserticola Kugler, 1978 PA: Palaearctic Acknowledgements Genus ROSSIMYIOPS Mesnil I would like to thank Dr. Amnon Freidberg and Dr. Tatyana Novoselsky, Tel Aviv University, Israel for supplying me with some relevant papers. I am grateful to Dr. Ashraf Al-Torky, Plant Protection Institute, Ministry of Agricul- – Rossimyiops achilleae (Kugler, 1972) ture, Egypt, for his help throughout this study. – Rossimyiops magnificus (Kugler, 1972) Funding No funding. Tribe SIPHONINI Genus ACTIA Robineau-Desvoidy Availability of data and materials Data supporting the conclusions of this article are presented in the main manuscript. – Actia crassicornis (Meigen, 1824) Author’s contributions This is a single author study. The author read and approved the final Genus PERIBAEA Robineau-Desvoidy manuscript. Ethics approval and consent to participate – Peribaea orbata (Wiedemann, 1830) Not applicable. – Peribaea palaestina (Villeneuve, 1934) Consent for publication Not applicable. Genus SIPHONA Meigen Subgenus SIPHONA Meigen, 1803 Competing interests The author declares that he has no competing interests. – Siphona (Siphona) efflatouni Mesnil, 1960 Publisher’sNote Springer Nature remains neutral with regard to jurisdictional claims in Tribe TACHININI. published maps and institutional affiliations. Genus PELETERIA Robineau-Desvoidy. Received: 18 December 2017 Accepted: 19 March 2018 Peleteria meridionalis (Robineau-Desvoidy, 1830). Abbreviations References List of abbreviations used throughout the catalogue Becker T (1908) Dipteren der Kanarischen Inseln. Mitt Zool Mus Berlin 4:1–180 C.: Central; E.: East; Eur.: Europe; ICZN: International Commission on Zoological Becker T (1910) Dipteren aus Südarabien und von der Insel Sokótra. 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Egyptian Journal of Biological Pest ControlSpringer Journals

Published: May 30, 2018

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