Molecular Biology and Evolution (ISSN 0737-4038) is a peer-reviewed Postal information journal published twelve times a year by Oxford University Press for the Molecular Biology and Evolution (ISSN 0737-4038) is published MOLECULAR BIOLOGY AND EVOLUTION Society for Molecular Biology and Evolution. monthly by Oxford University Press, 2001 Evans Road, Cary, NC 27513-2009. Periodicals Postage Paid at Cary, NC, and additional Molecular Biology and Evolution (MBE) publishes research at the mailing ofﬁces. POSTMASTER: Send address changes to Molecular interface between molecular and evolutionary biology. The journal (ISSN 0737-4038) Biology and Evolution, Mercury Media Processing, 187 E. Albert publishes molecular evolutionary patterns, processes, and predictions at St., Rahway, NJ 07065, USA. all levels of taxonomic, functional, genomic, and phenotypic organization. It also publishes new and improved methods, resources, and theories Founding Editors: Walter Fitch and Masatoshi Nei Environmental and ethical policies that are critical for advancing the molecular evolutionary research. Editor in Chief: Sudhir Kumar Institute for Genomics and Oxford Journals, a division of Oxford University Press, is committed MBE does not publish manuscripts containing largely descriptive work, Evolutionary Medicine, to working with the global community to bring the highest quality conﬁ rmatory research, and discoveries with a limited scope or impact. research to the widest possible audience. Oxford Journals will protect Temple University, Membership in the Society for Molecular Biology and Evolution the environment by implementing environmentally friendly policies Philadelphia, PA, USA Individuals interested in becoming a member of the Society for and practices wherever possible. Please see academic.oup.com/journals/ EIC.MBE@gmail.com Molecular Biology and Evolution can obtain application forms from pages/authors_ethics for further information on environmental and email@example.com. ethical policies. Editorial Assistant: Marjorie Gillis EAssist.MBE@gmail.com Subscriptions Digital Object Identiﬁ ers Editor’s Advisors: Barry Hall firstname.lastname@example.org A subscription to Molecular Biology and Evolution comprises 12 issues. For information on dois and to resolve them, please visit www.doi.org. Marcy Uyenoyama email@example.com All prices include postage; for subscribers outside the Americas, issues are sent air freight. Molecular Biology and Evolution Advance Access News Writer: Joseph Caspermeyer firstname.lastname@example.org Permissions contains papers that have recently been accepted but have not yet been For information on how to request permissions to reproduce articles or Assistant Managing Editor: Heather Rowe email@example.com included within an issue. Advance Access is updated daily. information from this journal, please visit academic.oup.com/journals/ pages/authors Editors (Senior and Associate) Annual Subscription Rate (Volume 34, 12 issues, 2017) Institutional A list of senior and associate editors is available at Advertising Print edition and site-wide online access: US$1201.00/£686.00/E1071.00 academic.oup.com/mbe/pages/Editorial_Board Advertising, inserts, and artwork enquiries should be addressed to Print edition only: US$1103.00/£631.00/E985.00 Advertising and Special Sales, Oxford Journals, Oxford University Site-wide online access only: US$780.00/£445.00/E694.00 Press, Great Clarendon Street, Oxford, OX2 6DP, UK. Tel: +44 (0)1865 Information for Authors Please note: UK£ rate applies to UK and Rest of World, except US 354767; Fax: +44 (0)1865 353774; E-mail: firstname.lastname@example.org. Guidelines for author submissions are available at and Canada ($) and Europe (E). Payment for orders to be delivered academic.oup.com/mbe/pages/General_Author_Guidelines within Europe should be made in euros. Granting of License There are other subscription rates available; for a complete listing, It is a condition of publication in the journal that authors grant please visit academic.oup.com/mbe/subscribe an exclusive license to the Society for Molecular Biology and Full prepayment in the correct currency is required for all orders. Payment Evolution. This ensures that requests from third parties to reproduce should be in US dollars for orders being delivered to the USA or Canada; articles are handled efﬁciently and consistently and will also allow the SOCIETY FOR MOLECULAR BIOLOGY AND EVOLUTION Euros for orders being delivered within Europe (excluding the UK); GBP article to be as widely disseminated as possible. As part of the license www.smbe.org sterling for orders being delivered elsewhere (i.e. not being delivered to agreement, authors may use their own material in other publications USA, Canada, or Europe). All orders should be accompanied by full provided that the journal is acknowledged as the original place of COUNCIL payment and sent to your nearest Oxford Journals ofﬁce. Subscriptions publication and Oxford University Press as the publisher. are accepted for complete volumes only. Orders are regarded as ﬁrm, Laura Landweber, President Statements and opinions expressed in publications of the Society for and payments are not refundable. Our prices include Standard Air Department of Ecology & Evolutionary Biology, Princeton University, Princeton, NJ 08544. Email: lﬂ @princeton.edu Molecular Biology and Evolution or in presentations given during its as postage outside of the UK. Claims must be notiﬁed within four months William F. Martin, President-elect regular meetings are those of the author(s) and do not necessarily reﬂect of despatch/order date (whichever is later). Subscriptions in the EEC may Institut für Molekulare Evolution, Heinrich Heine-Universität, Düsseldorf, Germany. Email: email@example.com the ofﬁcial position of the Society for Molecular Biology and Evolution, be subject to European VAT. If registered, please supply details to avoid the Editors, or the organizations with which the authors are afﬁliated. George Zhang, Past President unnecessary charges. For subscriptions that include online versions, a The Editor(s), publisher, and Society disclaim any responsibility or Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109. Email: firstname.lastname@example.org proportion of the subscription price may be subject to UK VAT. liability for such material and do not guarantee, warrant, or endorse any Subscribers in Canada, please add GST to the prices quoted. Personal rate David Pollock, Secretary product or service mentioned, including those by advertisers, nor do they subscriptions are only available if payment is made by personal cheque or University of Colorado School of Medicine Biochemistry and Molecular Genetics, Aurora, CO 80045. Email: email@example.com guarantee any claim made by the manufacturer of such product or service. credit card, delivery is to a private address, and is for personal use only. Ofﬁcial positions of the Society are established only by its Council. Juliette de Meaux, Treasurer The current year and two previous years’ issues are available from Oxford Institute for Evolution and Biodiversity, University of Münster, Germany 48149. Email: firstname.lastname@example.org Copyright © 2017 Society for Molecular Biology and Evolution Journals. Previous volumes can be obtained from the Periodicals Service Joanna Masel, Councillor (2017–2019) Company at http://www.periodicals.com/oxford.html or Periodicals Service Department of Ecology & Evolutionary Biology, University of Arizona, Tucson, AZ 85721. Email: email@example.com All rights reserved; no part of this publication may be reproduced, Company, 11 Main Street, Germantown, NY 12526, USA. E-mail: stored in a retrieval system, or transmitted in any form or by any Jay Storz, Councillor (2017–2019) firstname.lastname@example.org. Tel: (518) 537-4700. Fax: (518) 537-5899. means, electronic, mechanical, photocopying, recording, or otherwise University of Nebraska School of Biological Sciences, Lincoln, NE 68339. Email: email@example.com Contact information without prior written permission of the publisher or a license Maud Tenaillon, Councillor (2015–2017) Journals Customer Service Department, Oxford University Press, Great permitting restricted copying issued in the UK by the Copyright Department Quantitative Genetics and Evolution, Le Moulon, 91190 Gif-sur-Yvette, France. Email: firstname.lastname@example.org Clarendon Street, Oxford OX2 6DP, UK. E-mail: jnls.cust. Licensing Agency Ltd, 90 Tottenham Court Road, London W1P 9HE. email@example.com. Tel: +44 (0)1865 353907. Fax: + 44 (0)1865 353485. Adam Eyre-Walker, Councillor (2015–2017) Typeset by Cenveo Publisher Services, Bengaluru, India; Printed by CTPS In the Americas, please contact: Journals Customer Service Department, School of Biological Sciences University of Sussex, Brighton BN1 9QG, United Kingdom. Email: firstname.lastname@example.org DIGIASIA Pte Ltd, Singapore. Oxford University Press, 2001 Evans Road, Cary, NC 27513, USA. Emma Teeling, Councillor (2015–2017) E-mail: email@example.com. Tel: (800) 852-7323 (toll-free in USA/ School of Biology and Environmental Science, University College Dublin, Dublin 4, Ireland. Email: Emma.Teeling@ucd.ie Indexing and Abstracting Canada) or (919) 677-0977. Fax: (919) 677-1714. In Japan, please Molecular Biology and Evolution is abstracted or indexed in Kateryna Makova, Councillor (2015–2017) contact: Journals Customer Service Department, Oxford University Biological Abstracts, BIOSIS, Chemical Abstracts, Current Contents Department of Biology, Penn State University, PA 16802. Email: firstname.lastname@example.org Press, 4-5-10-8F Shiba, Minato-ku, Tokyo, 108-8386, Japan. E-mail: in Life Sciences, Excerpta Medica, Index Medicus, ISI/BIOMED, email@example.com. Tel: (81) 3 5444 5858. Fax: (81) 3 3454 2929. Science Citation Index, and Zoological Record. Methods of payment (i) Check (payable to Oxford University Press, to Oxford University Oxford University Press is a department of the University of Oxford. It Press, Cashiers Ofﬁce, Great Clarendon Street, Oxford OX2 6DP, UK) furthers the University’s objective of excellence in research, scholarship, in GB£ Sterling (drawn on a UK bank), US$ Dollars (drawn on a US and education by publishing worldwide. bank), or EUE Euros. (ii) Bank transfer to Barclays Bank Plc, Oxford Group Ofﬁce, Oxford (bank sort code 20-65-18) (UK), overseas only Swift code BARC GB 22 (GB£ Sterling to account no. 70299332, IBAN GB89BARC20651870299332; US$ Dollars to account no. 66014600, IBAN GB27BARC20651866014600; EUE Euros to account no. 78923655, IBAN GB16BARC20651878923655). (iii) Credit card (Mastercard, Visa, Switch or American Express). JMOLBEV1701M m mbe_34_1_Cover.indd 2 be_34_1_Cover.indd 2 2 21-12-2016 12:44:30 1-12-2016 12:44:30
Molecular Biology and Evolution – Oxford University Press
Published: Jan 1, 2017
It’s your single place to instantly
discover and read the research
that matters to you.
Enjoy affordable access to
over 12 million articles from more than
10,000 peer-reviewed journals.
All for just $49/month
Read as many articles as you need. Full articles with original layout, charts and figures. Read online, from anywhere.
Keep up with your field with Personalized Recommendations and Follow Journals to get automatic updates.
It’s easy to organize your research with our built-in tools.
Read from thousands of the leading scholarly journals from SpringerNature, Elsevier, Wiley-Blackwell, Oxford University Press and more.
All the latest content is available, no embargo periods.
“Hi guys, I cannot tell you how much I love this resource. Incredible. I really believe you've hit the nail on the head with this site in regards to solving the research-purchase issue.”Daniel C.
“Whoa! It’s like Spotify but for academic articles.”@Phil_Robichaud
“I must say, @deepdyve is a fabulous solution to the independent researcher's problem of #access to #information.”@deepthiw
“My last article couldn't be possible without the platform @deepdyve that makes journal papers cheaper.”@JoseServera