Agriculture, Ecosystems and Environment 249 (2017) 31–37 Contents lists available at ScienceDirect Agriculture, Ecosystems and Environment journal homepage: www.elsevier.com/locate/agee Inﬂuence of farmland abandonment on the species composition of wetland MARK ground beetles in Kushiro, Japan a,b, c b,d b Satoshi Yamanaka , Takumi Akasaka , Yuki Yabuhara , Futoshi Nakamura Hokkaido Research Center, Forestry and Forest Products Research Institute, Hitsujigaoka 7, Sapporo 062-8516, Japan Department of Forest Science, Graduate School of Agriculture, Hokkaido University, Kitaku Kita9 Nishi9, Sapporo 060-8589, Japan Laboratory of Conservation Ecology, Obihiro University of Agriculture and Veterinary Medicine, Nishi 2-sen 11, Inadacho, Obihiro 080-8555, Japan Institute of Technology and Science, The University of Tokushima, 2-1 Minami-Josanjima, Tokushima 770-8586, Japan ARTICLE I NFO ABSTRACT Keywords: Depopulation trends in many developed regions are resulting in an increase in areas of abandoned farmland, Agricultural landscape which could provide an alternative habitat for species endangered by past conversion of wetlands for agriculture. Biodiversity Additionally, various spatial and temporal factors (landscape structure, local habitat quality, and abandonment Carabid age) could inﬂuence species composition in abandoned farmland. In this study, we explored the spatio-temporal Fen eﬀects of land abandonment on the species composition of wetland ground beetles (Coleoptera: Carabidae) to Passive restoration
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