1 <h5>Introduction</h5> A great deal of progress has been made in developing and refining techniques to identify representative systems of conservation areas- comprising both formal reserves and a variety of complementary off-reserve management approaches ( Pressey et al., 1993; Margules and Pressey, 2000; Possingham et al., 2000; Pressey and Cowling, 2001 ). However, in addition to representing biodiversity, a goal of establishing conservation areas should be the persistence of the species and processes they contain. This is achieved by excluding processes that threaten biodiversity ( Faith and Walker, 1996; Pressey et al., 1996 ) and by designing systems that support the population, ecological and evolutionary processes that maintain and generate biodiversity ( Smith et al., 1993; Balmford et al., 1998; Margules and Pressey, 2000 ). In addition to spatial considerations, achieving persistence requires the consideration of time over a wide range of scales. Surprisingly little attention has been given to many temporal phenomena in conservation planning. Perhaps the best developed aspect of temporal issues in planning has been the research and applications on population dynamics and persistence ( Soulé and Terborgh, 1999; Hanski and Ovaskainen, 2000; Reed et al., 2002 ). Other important processes have received much less
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