WRKY transcription factors ( TF s) are well known for regulating plant abiotic and biotic stress tolerance. However, much less is known about how WRKY TF s affect plant-specialized metabolism. Analysis of WRKY TF s regulating the production of specialized metabolites emphasizes the values of the family outside of traditionally accepted roles in stress tolerance. WRKYs with conserved roles across plant species seem to be essential in regulating specialized metabolism. Overall, the WRKY family plays an essential role in regulating the biosynthesis of important pharmaceutical, aromatherapy, biofuel, and industrial components, warranting considerable attention in the forthcoming years. Glossary TF transcription factor bHLH basic helix-loop-helix TIA terpene indole alkaloid BIA benzylisoquinoline alkaloid
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