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Animal Biology , Vol. 55, No. 4, pp. 297-321 (2005) Koninklijke Brill NV, Leiden, 2005. Also available online - www.brill.nl Facsimile of On aims and methods of Ethology NIKO TINBERGEN Department of Zoology, University of Oxford This paper was originally published as: Tinbergen, N. (1963) On aims and methods of ethology. Zeitschrift für Tierpsy- chologie , 20, 410-433. This journal was renamed Ethology in 1986. We are grateful to the current publishers and copyright-owners, Blackwell Publish- ing, Oxford, for their kind permission to reproduce this paper.298 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 299300 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 301302 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 303304 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 305306 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 307308 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 309310 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 311312 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 313314 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 315316 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 317318 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 319320 Facsimile of Tinbergen (1963)Facsimile of Tinbergen (1963) 321
Animal Biology – Brill
Published: Jan 1, 2005
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